Articles
Chelonian Conservation and Biology
International Journal of Turtle and Tortoise Research
(ISSN 1071-8443)
Volumes 1–4 (1993–2005)
Volume 1 (1993–1995)
Volume 1, Number 1 – November 1993
Prologue
Editorial: A new turtle and tortoise journal. JOHN L. BEHLER, PETER C.H. PRITCHARD, AND ANDERS G.J. RHODIN 2
Articles
Conservation biology of the pig-nosed turtle, Carettochelys insculpta. ARTHUR GEORGES AND MARK ROSE 3
Status and exploitation of the Madagascan big-headed turtle, Erymnochelys madagascariensis. GERALD KUCHLING AND RUSSELL A. MITTERMEIER 13
Distribution, osteology, and natural history of the Asian giant softshell turtle, Pelochelys bibroni, in Papua New Guinea. ANDERS G.J. RHODIN, RUSSELL A. MITTERMEIER, AND PHILIP M. HALL 19
Carapacial pankinesis in the Malayan softshell turtle, Dogania subplana. PETER C.H. PRITCHARD 31
On growth, sexual dimorphism, and the general ecology of the African spurred tortoise, Geochelone sulcata, in Mali. MICHAEL R.K. LAMBERT 37
Notes and Field Reports
Range extension for Emydura subglobosa in Papua New Guinea. ANDERS G.J. RHODIN 47
A ranching project for freshwater turtles in Costa Rica. PETER C.H. PRITCHARD 48
New localities for Pyxis planicauda in west-central Madagascar. JOHN L. BEHLER, QUENTIN M.C. BLOXAM, EDMOND R. RAKOTOVAO, AND HERILALA J.A.R. RANDRIAMAHAZO 49
Reunion Island – still a land of tortoises. JUAN GONZALEZ GONZALEZ 51
Translations
On the distribution, relative abundance and protection of tortoises in Bulgaria. VLADIMIR A. BESHKOV [1984]. Translation from Bulgarian arranged by JAMES R. BUSKIRK 53
Newsnotes and Announcements
International conference on conservation, restoration, and management of turtles and tortoises. MICHAEL W. KLEMENS, KRISTIN H. BERRY, AND SUZANNE DOHM 63
Bibliographic database of chelonian literature established. ANDERS G.J. RHODIN, KENT R. BEAMAN, AND LORI A. DICONZA 64
Chelonian Research Foundation Linnaeus Fund: 1992 grant recipients. ANDERS G.J. RHODIN 64
Appendix
IUCN/SSC Tortoise and Freshwater Turtle Specialist Group 1993 membership list. 67
Volume 1, Number 2 – July 1994
Prologue
Editorial comment. JOHN L. BEHLER, PETER C.H. PRITCHARD, AND ANDERS G.J. RHODIN 79
Articles
The Mary River tortoise: a new genus and species of short-necked chelid from Queensland, Australia (Testudines; Pleurodira). JOHN CANN AND JOHN M. LEGLER 81
Activity patterns and habitat use of box turtles (Terrapene carolina bauri) on a Florida island, with recommendations for management. C. KENNETH DODD, JR., RICHARD FRANZ, AND LORA L. SMITH 97
The ecology of sea beach nesting in slider turtles (Trachemys scripta venusta) from Caribbean Costa Rica. DON MOLL 107
Morphometric analysis and taxonomy of cooter and red-bellied turtles in the North American genus Pseudemys (Emydidae). MICHAEL E. SEIDEL 117
Comments on Chinemys palaeannamitica and certain other chelonian taxa based upon material from archaeological sites. PETER C.H. PRITCHARD 131
Notes and Field Reports
Husbandry and captive breeding of the parrot-beaked tortoise, Homopus areolatus. JAMES E. BARZYK 138
Thermal limits of incubation in embryos of softshell turtles (Apalone mutica). MICHAEL V. PLUMMER, CHRIS E. SHADRIX, AND REBEKAH C. COX 141
Reproduction of the cabeçudo, Peltocephalus dumerilianus, in the Biological Reserve of Rio Trombetas, Pará, Brazil. RICHARD C. VOGT, VITOR HUGO CANTARELLI, AND ALBERTO GUERREIRO DE CARVALHO 145
Maximum size and clutch size records for eastern painted turtles, Chrysemys picta picta, from mid-coastal Maine. ANDERS G.J. RHODlN AND GLEN H. MITTELHAUSER 148
Translations
Status and conservation prospects of Testudo graeca L. inhabiting the Black Sea coast of the Caucasus. ALEXANDER A. INOZEMTSEV AND SOLOMON L. PERESHKOLNlK. [1985]. Translation from Russian by MARINA PROUTKINA, arranged by DEWEY I. WALLACE 151
Commentaries and Reviews
Temperature controlled sex determination as a tool for turtle conservation. RICHARD C. VOGT 159
Newsnotes and Announcements
Turtle and tortoise papers presented at the Second World Congress of Herpetology. 163
Chelonia Neotropica – a new turtle and tortoise newsletter from Brazil. ALINE TRISTAO BERNARDES 165
Chelonian Research Foundation Linnaeus Fund: 1993 grant recipients. ANDERS G.J. RHODIN 166
Volume 1, Number 3 – January 1995
Prologue
Editorial comment. JOHN L. BEHLER, PETER C.H. PRITCHARD, AND ANDERS G.J. RHODIN 171
Articles
Movements and home range of hatchling and yearling gopher tortoises, Gopherus polyphemus. JOSEPH A. BUTLER, RAY D. BOWMAN, TODD W. HULL, AND SCOTT SOWELL 173
Reproductive biology of the chopontil, Claudius angustatus (Testudines: Kinosternidae), in southern Veracruz, Mexico. OSCAR A. FLORES-VILLELA AND GEORGE R. ZUG 181
Early post-emergent behavior and habitat selection in hatchling Blanding’s turtles, Emydoidea blandingii, in Massachusetts. BRIAN O. BUTLER AND TERRY E. GRAHAM 187
An assessment of the effects of commercial exploitation on the pancake tortoise, Malacochersus tornieri, in Tanzania. MICHAEL W. KLEMENS AND DON MOLL 197
Habitat use and population parameters of the spotted turtle, Clemmys guttata, a Species of Special Concern in Massachusetts. TERRY E. GRAHAM 207
Observations and morphometric data on the Namaqualand speckled tortoise, Homopus signatus signatus (Gmelin, 1789), in South Africa. NICOLAS BAYOFF 215
Notes and Field Reports
Exploitation of the alligator snapping turtle, Macroclemys temminckii, in Louisiana: a case study. KEVIN SLOAN AND JEFFREY E. LOVICH 221
Turtles at a market in western Yunnan: possible range extensions for some southern Asiatic chelonians in China and Myanmar. GERALD KUCHLING 223
Reproductive biology of the Indian roofed turtle, Kachuga tecta, in Bangladesh. MD. LOKMAN HOSSAIN AND MD. SOHRAB UDDIN SARKER 226
Incubation period and sex ratio of Hermann’s tortoise, Testudo hermanni boettgeri. BERT T. EENDEBAK 227
Growth of head-started Kemp’s ridley sea turtles (Lepidochelys kempii) following release. CHARLES W. CAILLOUET, JR., CLARK T. FONTAINE, SHARON A. MANZELLA-TIRPAK, AND THEODORE D. WILLIAMS 231
Translations
Population density of the steppe tortoise in some regions of the Almaty and Taldyqorghan Districts, Kazakhstan. RUDOLF A. KUBYKIN. [1988]. Translation from Russian by CHERYL A. WHlTE, arranged by JAMES R. BUSKIRK 235
Commentaries and Reviews
Manipulating sex ratios: turtle speed ahead! N. MROSOVSKY AND MATTHEW H. GODFREY 238
Workshop on the ecology, status, and management of the diamondback terrapin (Malaclemys terrapin), Savannah River Ecology Laboratory, 2 August 1994: final results and recommendations. Compiled by RICHARD A. SEIGEL AND J. WHITFIELD GIBBONS 240
Turtle issues at CITES 1994. PETER C.H. PRITCHARD 243
In Memoriam. Leo Brongersma — an appreciation. PETER C.H. PRITCHARD 244
Newsnotes and Announcements
Chelonian Research Foundation Linnaeus Fund: 1994 grant recipients. ANDERS G.J. RHODIN 248
Research activities of the Sea Turtle Research Unit (SEATRU) of Universiti Pertanian Malaysia. SEA TURTLE RESEARCH UNIT 248
Centre de Reproducció de Tortugues de l’Albera in Spain. RAMON MASCORT BRUGAROLAS 249
New publication on turtle and tortoise trade in the United States. ALLEN SALZBERG 250
Chelonian Research Monographs, a new publication series by Chelonian Research Foundation. ANDERS G.J. RHODIN 250
Volume 1, Number 4 – August 1995
Articles
On geographical size variation, growth, and sexual dimorphism of the leopard tortoise, Geochelone pardalis, in Somaliland. MICHAEL R.K. LAMBERT 269
Mycoplasmosis and the desert tortoise (Gopherus agassizii) in Las Vegas Valley, Nevada. ELLIOTT R. JACOBSON, MARY B. BROWN, ISABELLA M. SCHUMACHER, BOBBY R. COLLINS, RICHARD K. HARRIS, AND PAUL A. KLEIN 279
Survival of head-started Kemp’s ridley sea turtles (Lepidochelys kempii) released into the Gulf of Mexico or adjacent bays. CHARLES W. CAILLOUET, JR., CLARK T. FONTAINE, SHARON A. MANZELLA-TIRPAK, AND DONNA J. SHAVER 285
Geographic variation in the matamata turtle, Chelus fimbriatus, with observations on its shell morphology and morphometry. MARCELO R. SANCHEZ-VILLAGRA, PETER C.H. PRITCHARD, ALFRED PAOLILLO, AND OMAR J. LINARES 293
Redescription and neotype designation of Pelochelys bibroni from southern New Guinea (Testudines: Trionychidae). ROBERT G. WEBB 301
A new batagurid turtle from northern Sulawesi, Indonesia. WILLIAM P. MCCORD, JOHN B. IVERSON, AND BOEADI 311
Notes and Field Reports
Salvage of eggs from road-killed red-eared sliders, Trachemys scripta elegans. JOHN K. TUCKER 317
Repatriation of radiated tortoises, Geochelone radiata, from Reunion Island to Madagascar. STEPHANE BOULLAY 319
Observations on feeding habits of Hydromedusa maximiliani (Testudines: Chelidae) in southeastern Brazil. FRANCO LEANDRO SOUZA AND AUGUSTO SHINYA ABE 320
The date of publication of Gray’s Catalogue of Shield Reptiles. ROBERT G. WEBB 322
Translations
Results of studies on the chelonians of Albania. IDRIZ HAXHIU. [1985]. Translation from Albanian by JAMES R. BUSKIRK 324
Current data on the chelonians of Albania. IDRIZ HAXHIU 326
Commentaries and Reviews
A clarification on the activities of Projeto TAMAR, Brazil. CECILIA BAPTISTOTTE 328
Systematics of the Pseudemys concinna-floridana complex (Testudines: Emydidae): an alternative interpretation. DALE R. JACKSON 329
How many species of cooter turtles and where is the scientific evidence? – a reply to Jackson. MICHAEL E. SEIDEL 333
Book Review: ACUNA MESEN, R.A. 1993. Las Tortugas Continentales de Costa Rica. San Jose, Costa Rica: Editorial ICER, 58 pp. MAHMOOD SASA 336
Newsnotes and Announcements
Turtle and tortoise papers presented at the International Congress of Chelonian Conservation. 338
Volume 2 (1996–1997)
Volume 2, Number 1 – February 1996
Prologue
Editorial comment. JOHN L. BEHLER, PETER C.H. PRITCHARD, AND ANDERS G.J. RHODIN 3
Articles
Overwintering in the nest chamber by hatchling painted turtles, Chrysemys picta, in northern New Jersey. JOSEPH A. DEPARI 5
Reproductive biology of a flood-season nesting freshwater turtle of the northern neotropics: Dermatemys mawii in Belize. JOHN POLISAR 13
Ecological characteristics of the pancake tortoise, Malacochersus tornieri, in Tanzania. DON MOLL AND MICHAEL W. KLEMENS 26
Observations on ecological changes threatening a population of Testudo graeca graeca in the Souss Valley, southern Morocco. JANE R. BAYLEY AND ANDY C. HIGHFIELD 36
Seasonal variation of embryonic diapause in the striped mud turtle (Kinosternon baurii) and general considerations for conservation planning. MICHAEL A. EWERT AND DAWN S. WILSON 43
A six season study of marine turtle nesting at Praia do Forte, Bahia, Brazil, with implications for conservation and management. MARIA ANGELA MARCOVALDI AND ANTONIO LAURENT 55
A survey of organized turtle watch participants on sea-turtle nesting beaches in Florida. STEVE A. JOHNSON, KAREN A. BJORNDAL, AND ALAN B. BOLTEN 60
Observations on the natural history and exploitation of the turtles of Nepal, with life history notes on Melanochelys trijuga. JOSEPH C. MITCHELL AND ANDERS G.J. RHODIN 66
Notes and Field Reports
The origin of marine turtles: a pluralistic view of evolution. JOSEPH J. KINNEARY 73
Occurrence and diet of juvenile loggerhead sea turtles, Caretta caretta, in the northwestern Gulf of Mexico. PAMELA T. PLOTKIN 78
First record of Heosemys spinosa from the Philippines, with biogeographic notes. INDRANEIL DAS 80
Neural bones in Australian chelid turtles. SCOTT THOMSON AND ARTHUR GEORGES 82
Temperature-dependent sex determination and hatching success in the gopher tortoise (Gopherus polyphemus). RUSSELL L. BURKE, MICHAEL A. EWERT, JEFFREY B. McLEMORE, AND DALE R. JACKSON 86
Marine turtle nesting, nest predation, hatch frequency, and nesting seasonality on the Osa Peninsula, Costa Rica. DANA L. DRAKE 89
Nesting activity of the loggerhead turtle, Caretta caretta, on Fethiye Beach, Turkey, in 1994. IBRAHIM BARAN AND OGUZ TURKOZAN 93
Stomach contents of commercially harvested adult alligator snapping turtles, Macroclemys temminckii. KEVIN N. SLOAN, KURT A. BUHLMANN, AND JEFFREY E. LOVICH 96
Distribution of nesting sites of sea turtles in Okinawajima and adjacent islands of the central Ryukyus, Japan. AKIRA KIKUKAWA, NAOKI KAMEZAKI, KOICHI HIRATE, AND HIDETOSHI OTA 99
Mating behavior in captive alligator snapping turtles (Macroclemys temminckii). J. BRENT HARREL, NEIL H. DOUGLAS, MAURY M. HARDAWAY, AND R. DALE THOMAS 101
Meat on the move: diet of a predatory turtle, Deirochelys reticularia (Testudines: Emydidae). DALE R. JACKSON 105
Kemp’s ridley sea turtle (Lepidochelys kempii) tracked by satellite telemetry from Louisiana to nesting beach at Rancho Nuevo, Tamaulipas, Mexico. MAURICE L. RENAUD, JAMES A. CARPENTER, JO A. WILLIAMS, AND ANDRE M. LANDRY, JR. 108
Observations on Manouria impressa at Phu Luang Wildlife Sanctuary, northeastern Thailand. TANYA CHAN-ARD, KUMTHORN THIRAKHUPT, AND PETER PAUL VAN DIJK 109
Commentaries and Reviews
Possible demographic and ecologic consequences of sex ratio manipulation in turtles. JEFFREY E. LOVICH 114
On the absence of spur-thighed tortoises, Testudo graeca, from Egypt. JAMES R. BUSKIRK 118
Turtle Poetry
To a box turtle. JOHN UPDIKE 121
Newsnotes and Announcements
Chelonian Research Foundation Linnaeus Fund: 1995 grant recipients. ANDERS G.J. RHODIN 122
Volume 2, Number 2 – October 1996
Special Focus Issue
The Leatherback Turtle, Dermochelys coriacea
Prologue
Editorial comment. JOHN L. BEHLER, PETER C.H. PRITCHARD, AND ANDERS G.J. RHODIN 139
Articles
Nesting Populations and Conservation
Leatherback turtles (Dermochelys coriacea) on St. Croix, U.S. Virgin Islands: fifteen years of conservation. RALF H. BOULON, JR., PETER H. DUTTON, AND DONNA L. McDONALD 141
Use of PIT tags and photoidentification to revise remigration estimates of leatherback turtles (Dermochelys coriacea) nesting in St. Croix, U.S. Virgin Islands, 1979–1995. DONNA L. McDONALD AND PETER H. DUTTON 148
Nesting of the leatherback turtle (Dermochelys coriacea) in Tongaland, KwaZulu-Natal, South Africa, 1963–1995. GEORGE R. HUGHES 153
Leatherback turtle, Dermochelys coriacea, nesting and nest success at Tortuguero, Costa Rica, in 1990–1991. ALISON J. LESLIE, DAVID N. PENICK, JAMES R. SPOTILA, AND FRANK V. PALADINO 159
Leatherback turtle, Dermochelys coriacea, nesting at Tortuguero, Costa Rica, in 1995. CATHI L. CAMPBELL, CYNTHIA J. LAGUEUX, AND JEANNE A MORTIMER 169
Nesting leatherback turtles at Las Baulas National Park, Costa Rica. ANTHONY C. STEYERMARK, KRISTINA WILLIAMS, JAMES R. SPOTILA, FRANK V. PALADINO, DAVID C. ROSTAL, STEPHEN J. MORREALE, MARIA TERESA KOBERG, AND RANDALL ARAUZ 173
Biology and conservation of leatherback turtles, Dermochelys coriacea, at Playa Langosta, Costa Rica. ANNY CHAVES, GUSTAVO SERRANO, GUILLERMO MARIN, EUGENIA ARGUEDAS, ALEJANDRO JIMENEZ, AND JAMES R. SPOTILA 184
Subsistence hunting of leatherback turtles, Dermochelys coriacea, in the Kai Islands, Indonesia. ALEXIS SUAREZ AND CHRISTOPHER H. STARBIRD 190
Decline of the leatherback population in Terengganu, Malaysia, 1956–1995. ENG-HENG CHAN AND HOCK-CHARK LIEW 196
Leatherback turtles, Dermochelys coriacea, nesting in French Guiana, 1978–1995. MARC GIRONDOT AND JACQUES FRETEY 204
Worldwide population decline of Dermochelys coriacea: are leatherback turtles going extinct? JAMES R. SPOTILA, ARTHUR E. DUNHAM, ALISON J. LESLIE, ANTHONY C. STEYERMARK, PAMELA T. PLOTKIN, AND FRANK V. PALADINO 209
Biology and Morphology
Respiratory physiology of adult leatherback turtles (Dermochelys coriacea) while nesting on land. FRANK V. PALADINO, JAMES R. SPOTILA, MICHAEL P. O’CONNOR, AND ROBERT E. GATTEN, JR. 223
Reproductive physiology of nesting leatherback turtles (Dermochelys coriacea) at Las Baulas National Park, Costa Rica. DAVID C. ROSTAL, FRANK V. PALADINO, RHONDA M. PATTERSON, AND JAMES R. SPOTILA 230
Shallow water diving by leatherback turtles in the South China Sea. SCOTT A. ECKERT, HOCK-CHARK LIEW, KAREN L. ECKERT, AND ENG-HENG CHAN 237
Age and growth in leatherback turtles, Dermochelys coriacea (Testudines: Dermochelyidae): a skeletochronological analysis. GEORGE R. ZUG AND JAMES F. PARHAM 244
Electron microscopic analysis of vascular cartilage canals in the humeral epiphysis of hatchling leatherback turtles, Dermochelys coriacea. JOHANNES A.G. RHODIN, ANDERS G.J. RHODIN, AND JAMES R. SPOTILA 250
Systematics and Evolution
On the availability of the name Dermochelys coriacea schlegelii (Garman, 1884) as a species or subspecies of the leatherback turtle. LEO DANIEL BRONGERSMA 261
Evolution and phylogeny of the leatherback turtles (Dermochelyidae), with descriptions of new fossil taxa. ROGER CONANT WOOD, JONNIE JOHNSON-GOVE, EUGENE S. GAFFNEY, AND KEVIN F. MALEY 266
Translations
Books on Marine Fish, in which True Figures of the Fish are Presented. Book XVI. Chapters II–V. On Turtles. GUILLAUME RONDELET. [1554]. Translated and annotated from Latin by KENNETH F. KITCHELL, JR., arranged and edited by ANDERS G.J. RHODIN 287
Commentaries and Reviews
Are leatherbacks really threatened with extinction? PETER C.H. PRITCHARD 303
Turtle Poetry
Awakening. KEVIN GRIMES 306
Volume 2, Number 3 – April 1997
Prologue
Editorial comment. ANDERS G.J. RHODIN, PETER C.H. PRITCHARD, JOHN L. BEHLER, AND RUSSELL A. MITTERMEIER 327
Articles
A new species of mud turtle (genus Kinosternon) from Jalisco and Colima, Mexico, with notes on its natural history. JAMES F. BERRY, MICHAEL E. SEIDEL, AND JOHN B. IVERSON 329
Life history and ecology of the Sonoran mud turtle (Kinosternon sonoriense) in southeastern Arizona: a preliminary report. RICHARD C. VAN LOBEN SELS, JUSTIN D. CONGDON, AND JOSIAH T. AUSTIN 338
Response of embryos of the red-eared turtle (Trachemys scripta elegans) to experimental exposure to water-saturated substrates. JOHN K. TUCKER, FREDRIC J. JANZEN, AND GARY L. PAUKSTIS 345
Growth, reproduction, and survivorship in the red-eared turtle, Trachemys scripta elegans, in Illinois, with conservation implications. JOHN K. TUCKER AND DON MOLL 352
Home range characteristics of radiotagged gopher tortoises on Kennedy Space Center, Florida. REBECCA B. SMITH, DAVID R. BREININGER, AND VICKIE L. LARSON 358
Ecology of the eastern spiny softshell, Apalone spinifera spinifera, in the Lamoille River, Vermont. TERRY E. GRAHAM AND ANDREW A. GRAHAM 363
Clutch size and frequency in Florida box turtles (Terrapene carolina bauri): implications for conservation. C. KENNETH DODD, JR. 370
A comparative spotting-scope study of the distribution and relative abundance of river cooters (Pseudemys concinna) in western Kentucky and southern Mississippi. PETER V. LINDEMAN 378
Redescription of the Arakan forest turtle Geoemyda depressa Anderson 1875 (Testudines: Bataguridae). JOHN B. IVERSON AND WILLIAM P. McCORD 384
Variation in shell morphology and color of Hermann’s tortoise, Testudo hermanni, in southern Europe. GHISLAINE GUYOT AND BERNARD DEVAUX 390
Young desert tortoise survival: energy, water, and food requirements in the field. KENNETH A. NAGY, DAVID J. MORAFKA, AND REBECCA A. YATES 396
Notes and Field Reports
The spotted turtle (Clemmys guttata) in central Florida. MARY E. BARNWELL, PETER A. MEYLAN, AND TIMOTHY WALSH 405
Radiographic determination of fecundity: is the technique safe for developing turtle embryos? THOMAS G. HINTON, PETER D. FLEDDERMAN, JEFFREY E. LOVICH, JUSTIN D. CONGDON, AND J. WHITFIELD GIBBONS 409
Maximum size of the Egyptian tortoise, Testudo kleinmanni. BALAZS L. FARKAS, LASZLO SASVARI, AND JAMES R. BUSKIRK 415
New locality records for freshwater turtles from the Yucatan Peninsula, Mexico. JAMES R. BUSKIRK 415
Problems for conservation of pond turtles (Emys orbicularis) in central Italy: is the introduced red-eared turtle (Trachemys scripta) a serious threat? LUCA LUISELLI, MASSIMO CAPULA, DARIO CAPIZZI, ERNESTO FILIPPI, VERONICA TRUJILLO JESUS, AND CLAUDIO ANIBALDI 417
Occurrence of double clutching in the southwestern pond turtle, Clemmys marmorata pallida, in the Los Angeles Basin. ROBERT H. GOODMAN, JR. 419
Allometry in the loggerhead turtle, Caretta caretta. NAOKl KAMEZAKl AND MASAFUMI MATSUI 421
Incidental capture of sea turtles in shrimp trawls with and without TEDs in U.S. Atlantic and Gulf waters. MAURICE L. RENAUD, JAMES M. NANCE, ELIZABETH SCOTT-DENTON, AND GREGG R. GITSCHLAG 425
Thermoregulation of the European pond turtle, Emys orbicularis, in central Italy. CORINNE DI TRANI AND MARCO A.L. ZUFFI 428
Dietary overlap in three sympatric congeneric freshwater turtles (Pseudemys) in Florida. KAREN A. BJORNDAL, ALAN B. BOLTEN, CYNTHIA J. LAGUEUX, AND DALE R. JACKSON 430
Predation by the imported fire ant (Solenopsis invicta) on loggerhead sea turtle (Caretta caretta) nests on Wassaw National Wildlife Refuge, Georgia. ROBERT A. MOULIS 433
Green turtle, Chelonia mydas, with fused carapacial scutes nesting at Atol das Rocas, Brazil. CLAUDIO BELLINI AND TAISI MARIA SANCHES 437
Commentaries and Reviews
Movement of hawksbill turtles — a different perspective on the DNA data. N. MROSOVSKY 438
Movement of hawksbill turtles: what scale is relevant to conservation, and what scale is resolvable with mtDNA data? BRIAN W. BOWEN AND ANNA L. BASS 440
Resolutions of the participants at the 17th Annual Symposium on Sea Turtle Biology and Conservation. Compiled by JEANETTE WYNEKEN AND DEBORAH T. CROUSE 442
Linnaeus Fund Research Reports
Seasonal movements and habitat preferences of spotted turtles (Clemmys guttata) in north central Connecticut. KATHLEEN M. PERILLO 445
Ecology and natural history of the wood turtle (Clemmys insculpta) in southern New Hampshire. SHEILA E. TUTTLE AND DAVID M. CARROLL 447
Geographic variation in the giant softshell turtle, Pelochelys bibroni. ROBERT G. WEBB 450
Orientation by the Australian eastern long-necked turtle, Chelodina longicollis. TERRY E. GRAHAM 450
Impact of egg content on post-hatching size, body composition, and performance in the common snapping turtle (Chelydra serpentina). MICHAEL S. FINKLER 452
Turtle Poetry
Ancient ones. GRACE S. McLAUGHLIN 456
Newsnotes and Announcements
Chelonian Research Foundation Linnaeus Fund: 1996 grant recipients. ANDERS G.J. RHODIN 457
The 17th Annual Sea Turtle Symposium — largest ever. JEANETTE WYNEKEN 457
El Mundo de la Tortuga: the leatherback turtle museum. CORINNE VIGNON, UBALDO ESTEBAN, LAURENT LOMBART, AND MAXIME MARQUEZ 458
The plight of the giant tortoises of Changuu Island, Zanzibar. STEVE TOLAN AND JOHN E. COOPER 459
Amphibian and Reptile Conservation — a new journal. CRAIG HASSAPAKIS 460
Three turtle conservation groups win Getty Prize. GILLIAN HAGGERTY 460
The 5th Annual Workshop on Natural History and Conservation of Florida Turtles. GEORGE L. HEINRICH 461
Volume 2, Number 4 – December 1997
Prologue
Editorial comment. Anders G.J. Rhodin, Peter C.H. Pritchard, John L. Behler, and Russell A. Mittermeier 481
Articles
Population structure, activity, and sexual dimorphism in a central Florida population of box turtles, Terrapene carolina bauri. Melissa A. Pilgrim, Terence M. Farrell, and Peter G. May 483
Behavior, hibernacula, and thermal relations of softshell turtles (Trionyx spiniferus) overwintering in a small stream. Michael V. Plummer and J. Cary Burnley 489
Home range of the turtle, Phrynops rufipes, in an isolated reserve in central Amazônia, Brazil. William E. Magnusson, Aldeniza Cardoso de Lima, Valdenise Lopes da Costa, and Richard C. Vogt 494
Restoration of epidermal scute patterns during regeneration of the chelonian carapace. Gerald Kuchling 500
Patterns of size and longevity of gopher tortoise (Gopherus polyphemus) burrows: implications for the longleaf pine ecosystem. Craig Guyer and Sharon M. Hermann 507
Activity, habitat, and movement patterns of softshell turtles (Trionyx spiniferus) in a small stream. Michael V. Plummer, Nathan E. Mills, and Steve L. Allen 514
Population structure, activity, and conservation of the neotropical freshwater turtle, Hydromedusa maximiliani, in Brazil. Franco Leandro Souza and Augusto Shinya Abe 521
Osteological characters of the shell and humerus in hinged tortoises of the African genus Kinixys. Donald G. Broadley 526
Age and growth of wild Kemp’s ridley turtles (Lepidochelys kempi): cumulative results of tagging studies in Florida. Jeffrey R. Schmid and Wayne N. Witzell 532
Nesting physiology of Kemp’s ridley sea turtles, Lepidochelys kempi, at Rancho Nuevo, Tamaulipas, Mexico, with observations on population estimates. David C. Rostal, Janice S. Grumbles, Richard A. Byles, René Márquez-M., and David Wm. Owens 538
The phylogeny of cheloniid sea turtles revisited. James F. Parham and David E. Fastovsky 548
Mauremys pritchardi, a new batagurid turtle from Myanmar and Yunnan, China. William P. McCord 555
Notes and Field Reports
Differences in body size among populations of red-eared sliders (Trachemys scripta elegans) subjected to different levels of harvesting. Lisa M. Close and Richard A. Seigel 563
Ecological distribution and population dynamics of the Mediterranean tortoise, Testudo graeca nikolskii, in xerophytic forests of the western Caucasus, southern Russia. Alexander A. Inozemtsev 567
Delayed feeding in neonatal Kemp’s ridley, Lepidochelys kempii: a captive sea turtle management technique. Clark T. Fontaine and Theodore D. Williams 573
Growth of the turtle, Phrynops rufipes, in central Amazônia, Brazil. William E. Magnusson, Aldeniza Cardoso de Lima, Valdenise Lopes da Costa, and Odilon Pimentel de Lima 576
Estimating the time between hatching of sea turtles and their emergence from the nest. Matthew H. Godfrey and N. Mrosovsky 581
Predation upon olive ridley sea turtles (Lepidochelys olivacea) by the American crocodile (Crocodylus acutus) at Playa Nancite, Costa Rica. Rudy M. Ortiz, Pamela T. Plotkin, and David Wm. Owens 585
Growth and reproductive estimates from alligator snapping turtles, Macroclemys temminckii, taken by commercial harvest in Louisiana. Anton D. Tucker and Kevin N. Sloan 587
Record-sized tortoises, Testudo graeca ibera and Testudo hermanni boettgeri, from Bulgaria. Vladimir A. Beshkov 593
Reproductive biology and demography of gopher tortoises (Gopherus polyphemus) from the western portion of their range. Kelley R. Smith, Janie A. Hurley, and Richard A. Seigel 596
Decline of the loggerhead turtle, Caretta caretta, nesting on Senri Beach in Minabe, Wakayama, Japan. Katsufumi Sato, Takeharu Bando, Yoshimasa Matsuzawa, Hideji Tanaka, Wataru Sakamoto, Shingo Minamikawa, and Kiyoshi Goto 600
A comparison of plastral scute lengths among members of the box turtle genera Cuora and Terrapene. Carl H. Ernst, Jeffrey E. Lovich, Arndt F. Laemmerzahl, and Steve Sekscienski 603
An ecotourism initiative to increase awareness and protection of marine turtles in Brazil: the Turtle by Night Program. Claudia Figueiredo Vieitas and Maria Ângela Marcovaldi 607
An anomalous specimen of Pelusios sinuatus lacking mesoplastra. Donald G. Broadley 610
Characteristics of hibernacula use by spotted turtles, Clemmys guttata, in Ohio. Timothy L. Lewis and John Ritzenthaler 611
Observations of reproductive behavior of male green turtles (Chelonia mydas) at a nesting beach in Cyprus. Annette C. Broderick and Brendan J. Godley 615
Kachuga sylhetensis recorded from northern Bengal, with notes on turtles of Gorumara National Park, eastern India. Indraneil Das 616
Commentaries and Reviews
Galápagos tortoise nomenclature: still unresolved. George R. Zug 618
Galápagos tortoise nomenclature: a reply. Peter C.H. Pritchard 619
In Memoriam. A man to envy: James J. Parsons, 1915–1997. David R. Stoddart 621
Linnaeus Fund Research Reports
Molecular phylogeography of the western pond turtle (Clemmys marmorata): preliminary results. Fredric J. Janzen, Stacia L. Hoover, and H. Bradley Shaffer 623
Translations
Reptiles of the Autonomous Republic of Turkmenistan. Chary A. Ataev. [1985]. Translated from Russian by Balázs Farkas, arranged by James R. Buskirk 627
Turtle Poetry
Wake, Methuselah! (Tortoise in midwinter at the London Zoo). Author Unknown 635
Newsnotes and Announcements
Chelonian conservation news. Allen Salzberg 636
Seychelles Giant Tortoise Conservation Project 638
Sea Turtle Survival League 638
Reviewers for Chelonian Conservation and Biology Volume 2 (1996 – 1997) 639
International Conference on Turtles and Tortoises 640
18th International Symposium on Sea Turtle Biology and Conservation 640
Volume 3 (1998–2000)
Volume 3, Number 1 – August 1998
Articles
Taxonomy of Indian Ocean giant tortoises (Dipsochelys). Justin Gerlach and Laura Canning 3
Distribution and habitats of the Euphrates softshell turtle, Rafetus euphraticus, in southeastern Anatolia, Turkey, with observations on biology and factors endangering its survival. Ertan Taskavak and Mehmet K. Atatür 20
Temperature effect on incubation period in the yellow-spotted river turtle, Podocnemis unifilis, in the Colombian Amazon. Vivian P. Páez and Brian C. Bock 31
Impact of agricultural development on a population of wood turtles (Clemmys insculpta) in southern Québec, Canada. Raymond A. Saumure and J. Roger Bider 37
On growth and morphometrics of leopard tortoises, Geochelone pardalis, in Serengeti National Park, Tanzania, with observations on effects of bushfires and latitudinal variation in populations of eastern Africa. Michael R.K. Lambert, Kenneth L.I. Campbell, and Jonathan D. Kabigumila 46
Decline of the olive ridley sea turtle (Lepidochelys olivacea) nesting assemblage at Nancite Beach, Santa Rosa National Park, Costa Rica. Roldán A. Valverde, Stephen E. Cornelius, and Claudette L. Mo 58
Population structure and movement patterns of alligator snapping turtles (Macroclemys temminckii) in northeastern Arkansas. Stanley E. Trauth, J.D. Wilhide, and Anthony Holt 64
Water transport by nesting painted turtles (Chrysemys picta marginata) in Michigan. Owen M. Kinney, Roy D. Nagle, and Justin D. Congdon 71
Demography and population status of the flattened musk turtle, Sternotherus depressus, in the Black Warrior River Basin of Alabama. Karan A. Bailey and Craig Guyer 77
Notes and Field Reports
Assessment and initial treatment of cold-stunned sea turtles. Samuel S. Sadove, Robert Pisciotta, and Robert DiGiovanni 84
Ecological notes on feral populations of Trachemys scripta elegans in northern Taiwan. Tien-Hsi Chen and Kuang-Yang Lue 87
Testosterone titers of immature loggerhead sea turtles (Caretta caretta) incidentally caught in the central Mediterranean: a preliminary sex ratio study. Paolo Casale, Guido Gerosa, Roberto Argano, Salvatore Barbaro, and Gaetano Fontana 90
Female-biased sex ratio of juvenile loggerhead sea turtles in Georgia. C. Robert Shoop, Carol A. Ruckdeschel, and Robert D. Kenney 93
Reproductive ecology of the Indian softshell turtle, Aspideretes gangeticus, in northern India. Karthikeyan Vasudevan 96
Morphometrics of the fossil box turtle, Terrapene innoxia Hay 1916, from Florida. Carl H. Ernst, James C. Wilgenbusch, Timothy P. Boucher, and Steven W. Sekscienski 99
Use of passive integrated transponder (PIT) tags for marking small freshwater turtles. Kurt A. Buhlmann and Tracey D. Tuberville 102
First record of fibropapilloma on an olive ridley turtle in Nicaragua. Todd M. Steiner, Randall Arauz Vargas, and Pandora Martinez 105
Food habits of the pelomedusid turtle Pelusios castaneus castaneus in southeastern Nigeria. Luca Luiselli 106
Annual deposition of scute rings in the western pond turtle, Clemmys marmorata. R. Bruce Bury and David J. Germano 108
Bait preferences of southeastern United States Coastal Plain riverine turtles: fish or fowl? John B. Jensen 109
Sexual dimorphism of neonate eastern spiny softshells, Apalone spinifera spinifera. Terry E. Graham and Cynthia B. Cobb 111
Molecules, morphology, and mud turtle phylogenetics (family Kinosternidae). John B. Iverson 113
Commentaries and Reviews
How to minimize risk and optimize information gain in assessing reproductive condition and fecundity of live female chelonians. Gerald Kuchling 118
Age determination in turtles: evidence of annual deposition of scute rings. David J. Germano and R. Bruce Bury 123
Linnaeus Fund Research Reports
Identification of Seychelles giant tortoises. Justin Gerlach and Laura Canning 133
Current status and conservation of the river cooter (Pseudemys concinna) in southern Illinois. Michael J. Dreslik 135
Of deadwood and map turtles (Graptemys): an analysis of species status for five species in three river drainages using replicated spotting-scope counts of basking turtles. Peter V. Lindeman 137
Nesting ecology of the striped mud turtle (Kinosternon baurii) in central Florida. Dawn S. Wilson 142
Morphology and histology of the digestive system of the desert tortoise, Gopherus agassizi. Robert M. Winokur, Martin Hicks, and Naomi Chaney 144
Turtle Poetry
Turtle Soup. Marilyn Chin 146
Newsnotes and Announcements
Chelonian conservation news. Allen Salzberg 147
Chelonian Research Foundation Linnaeus Fund: 1997 grant recipients. Anders G.J. Rhodin 150
Turtle decoys – a request for information. René E. Honegger 151
The 18th Annual Sea Turtle Symposium – Student Paper Awards. Anders G.J. Rhodin and F. Alberto Abreu Grobois 152
Symposium on the Status and Conservation of Florida Turtles. George L. Heinrich 152
Volume 3, Number 2 – April 1999
Special Focus Issue
The Hawksbill Turtle, Eretmochelys imbricata
Prologue
Editorial comment. Anders G.J. Rhodin and Peter C.H. Pritchard 171
Reviews
Biology and Status of the Hawksbill in the Caribbean.
A Review Prepared by the IUCN/SSC Marine Turtle Specialist Group.
Foreword. Maria Angela (Neca) Marcovaldi 173
Introduction. Conservation of hawksbill sea turtles: perceptions and realities. Karen A. Bjorndal 174
Status of the hawksbill turtle (Eretmochelys imbricata) in the Caribbean region. Anne B. Meylan 177
Population modeling and implications for Caribbean hawksbill sea turtle management. Deborah T. Crouse 185
International movements of immature and adult hawksbill turtles (Eretmochelys imbricata) in the Caribbean region. Anne B. Meylan 189
Genetic analysis to elucidate the natural history and behavior of hawksbill turtles (Eretmochelys imbricata) in the wider Caribbean: a review and re-analysis. Anna L. Bass 195
Articles
Status justification for listing the hawksbill turtle (Eretmochelys imbricata) as Critically Endangered on the 1996 IUCN Red List of Threatened Animals. Anne B. Meylan and Marydele Donnelly 200
Differential tag retention in Caribbean hawksbill turtles. Robert P. van Dam and Carlos E. Diez 225
Population structure of hawksbill turtles on a foraging ground in the Dominican Republic. Yolanda M. León and Carlos E. Diez 230
Internesting movements and behavior of hawksbill turtles (Eretmochelys imbricata) around Buck Island Reef National Monument, St. Croix, U.S. Virgin Islands. Christopher H. Starbird, Zandy Hillis-Starr, James T. Harvey, and Scott A. Eckert 237
Population ecology and demographic implications drawn from an 11-year study of nesting hawksbill turtles, Eretmochelys imbricata, at Jumby Bay, Long Island, Antigua, West Indies. James I. Richardson, Rebecca Bell, and Thelma H. Richardson 244
Estimating the annual size of hawksbill (Eretmochelys imbricata) nesting populations from mark-recapture studies: the use of long-term data to provide statistics for optimizing survey effort. Rhema Kerr, James I. Richardson, and Thelma H. Richardson 251
Reproduction and nesting of the hawksbill turtle, Eretmochelys imbricata, in the Cuban archipelago. Felix Moncada, Elvira Carrillo, Ana Saenz, and Gonzalo Nodarse 257
Hawksbill turtles (Eretmochelys imbricata) in Cuba: an assessment of the historical harvest and its impacts. Elvira Carrillo, Grahame J.W. Webb, and S. Charlie Manolis 264
Nesting of the hawksbill turtle, Eretmochelys imbricata, in Río Lagartos, Yucatán, Mexico, 1990–1997. Mauricio Garduño-Andrade 281
Increases in hawksbill turtle (Eretmochelys imbricata) nestings in the Yucatán Peninsula, Mexico, 1977–1996: data in support of successful conservation? Mauricio Garduño-Andrade, Vicente Guzmán, Emma Miranda, Raquel Briseño-Dueñas, and F. Alberto Abreu-Grobois 286
Genetic sourcing for the hawksbill turtle, Eretmochelys imbricata, in the northern Caribbean region. Rogelio Díaz-Fernández, Toshinao Okayama, Tomouki Uchiyama, Elvira Carrillo, Georgina Espinosa, René Márquez, Carlos Diez, and Hiroko Koike 296
Nesting and conservation management of hawksbill turtles (Eretmochelys imbricata) in northern Bahia, Brazil. Maria Angela Marcovaldi, Claudia F. Vieitas, and Matthew H. Godfrey 301
Juvenile Eretmochelys imbricata and Chelonia mydas in the Archipelago of Fernando de Noronha, Brazil. Taisi Maria Sanches and Claudio Bellini 308
The hawksbill turtle, Eretmochelys imbricata, in the Arabian Gulf. Nicolas J. Pilcher 312
Temporal distribution and periodicity in hawksbill turtles (Eretmochelys imbricata) nesting at Cousin Island, Republic of Seychelles, 1971–1997. Jeanne A. Mortimer and Roby Bresson 318
Hawksbill turtles, Eretmochelys imbricata, nesting on Redang Island, Terengganu, Malaysia, from 1993 to 1997. Eng-Heng Chan and Hock-Chark Liew 326
Reproductive biology of the hawksbill turtle, Eretmochelys imbricata, in Sabah, Malaysia. Nicolas J. Pilcher and Lamri Ali 330
Current status of nesting populations of the hawksbill turtle (Eretmochelys imbricata) in the Java Sea, Indonesia. Hiroyuki Suganuma, Naoki Kamezaki, and Akil Yusuf 337
Hawksbill turtle, Eretmochelys imbricata, nesting at Milman Island, northern Great Barrier Reef, Australia. Kirstin A. Dobbs, Jeffrey D. Miller, Colin J. Limpus, and André M. Landry, Jr. 344
Genetic diversity of the hawksbill turtle in the Indo-Pacific and Caribbean regions. Toshinao Okayama, Rogelio Díaz-Fernández, Yoshiyuki Baba, Matheus Halim, Osamu Abe, Naofumi Azeno, and Hiroko Koike 362
Turtle Poetry
Adelita sleeps…. Barbara Garrison 368
Caretta’s Return. Barbara Garrison 369
Newsnotes and Announcements
Chelonian conservation news. Allen Salzberg 370
Chelonian Research Foundation Linnaeus Fund: 1998 grant recipients. Anders G.J. Rhodin 373
The 19th Annual Sea Turtle Symposium – student presentation awards. Anders G.J. Rhodin and David Wm. Owens 374
2nd ASEAN Symposium and Workshop on Sea Turtle Biology and Conservation. Nicolas J. Pilcher 375
2nd International Symposium on Emys orbicularis. Alain Veysset and Barbara Livoreil 375
A Forum on the Biology and Conservation of Turtles in the Genus Clemmys. Sheila Tuttle 376
World’s first turtle shell stockpile to go up in flames as Miss World 1998 contestants look on. Jeanne A. Mortimer 376
New data reverse turtle’s spot on reptile tree. Carol Kaesuk Yoon 377
Sri Lanka herpetological publications available. Anslem de Silva 378
Chelonian Research Foundation announces a new turtle and tortoise newsletter and web site: www.chelonian.org. Anders G.J. Rhodin 378
Volume 3, Number 3 – December 1999
Prologue
Editorial comment. Anders G.J. Rhodin 391
Articles
Home range and microhabitat use in the angonoka (Geochelone yniphora) in Madagascar. Lora L. Smith, Bourou Robert, Mahatoly Joby, and Sibo Clément 393
Environmental correlates of hatchling emergence in the red-eared turtle, Trachemys scripta elegans, in Illinois. John K. Tucker 401
Thermal profiles of sea turtle hatcheries and nesting areas at Praia do Forte, Brazil. Eugênia Naro-Maciel, N. Mrosovsky, and Maria Ângela Marcovaldi 407
Ecology and status of a wood turtle (Clemmys insculpta) population in West Virginia. Allan J. Niederberger and Michael E. Seidel 414
Nesting ecology of Podocnemis sextuberculata (Testudines, Pelomedusidae) in the Japurá River, Amazonas, Brazil. Juarez C.B. Pezzuti and Richard C. Vogt 419
Habitat selection and movements of diamondback terrapins, Malaclemys terrapin, in a Maryland estuary. Willem M. Roosenburg, Katherin L. Haley, and Scott McGuire 425
Home ranges of spotted turtles (Clemmys guttata) in southwestern Ohio. Timothy L. Lewis and Craig A. Faulhaber 430
Leopard tortoises (Geochelone pardalis) in Valley Bushveld, Eastern Cape, South Africa: specialist or generalist herbivores? Mervyn C. Mason, Graham I.H. Kerley, Craig A. Weatherby, and William R. Branch 435
Occurrence and trade of the golden turtle, Cuora trifasciata, in Laos. Robert J. Timmins and Khamkhoun Khounboline 441
Physical and microhabitat characteristics of burrows used by juvenile desert tortoises (Gopherus agassizii). Dawn S. Wilson, C. Richard Tracy, Kenneth A. Nagy, and David J. Morafka 448
Conservation genetics of the giant Amazon River turtle (Podocnemis expansa; Pelomedusidae) – inferences from two classes of molecular markers. Jack W. Sites, Jr., Nancy N. FitzSimmons, Nelson Jorge da Silva, Jr., and Vitor H. Cantarelli 454
Notes and Field Reports
Growth of slider turtles (Trachemys scripta) from temperate and tropical populations. John K. Tucker, Don Moll, and Edward O. Moll 464
Husbandry, behavior, and captive breeding of the Namaqualand speckled padloper (Homopus signatus signatus). Victor J.T. Loehr 468
Diet of Elseya purvisi and Elseya georgesi (Testudines: Chelidae), a sibling species pair of freshwater turtles from eastern Australia. Mathew Allanson and Arthur Georges 473
Short-term effectiveness of Passive Integrated Transponder (PIT) tags used in the study of Mediterranean marine turtles. Brendan J. Godley, Annette C. Broderick, and Sinead Moraghan 477
On the turtles of Hainan Island, southern China. Ron W.F. de Bruin and Harald G. Artner 479
Observations on predation of marine turtle nests at Akyatan, Turkey, eastern Mediterranean. Monica Aureggi, Guido Gerosa, and Sedat V. Yerli 487
Occurrence of the furrowed wood turtle (Rhinoclemmys areolata) on Turneffe Atoll, Belize. Steven G. Platt, William B. Karesh, John B. Thorbjarnarson, and Thomas R. Rainwater 490
Size differences in hind limbs and carapaces of hatchling green turtles (Chelonia mydas) from Hawaii and Florida, USA. Jeanette Wyneken, George H. Balazs, S.K.K. Murakawa, and Yvette Anderson 491
Feeding behavior and the saddleback shell of Dipsochelys arnoldi. Justin Gerlach 496
Long-term trends in size of stranded juvenile loggerhead sea turtles (Caretta caretta). C. Robert Shoop, Carol A. Ruckdeschel, and Robert D. Kenney 501
Effects of short-term water temperature variation on food consumption in painted turtles (Chrysemys picta marginata). Kristie D. Gianopulos and John W. Rowe 504
Habitat structures associated with juvenile gopher tortoise burrows on pine plantations in Alabama. Matthew J. Aresco 507
First record of the Nile soft-shelled turtle, Trionyx triunguis, from Kos Island, Greece, with comments on its occurrence in the eastern Mediterranean. Ertan Taskavak, Michael J. Reimann, and Wouter N. Polder 510
Cutaneous surface area in freshwater turtles. Paul A. Stone and John B. Iverson 512
Stomach contents of an olive ridley turtle (Lepidochelys olivacea) from the Gulf of Papua, Papua New Guinea. C. Sylvia Spring and Janet Gwyther 516
First record of introduced eastern Mediterranean tortoises, Testudo hermanni boettgeri, in southern France. Ghislaine Guyot and Peter C.H. Pritchard 518
Commentaries and Reviews
Estimating the time between hatching and emergence from the nest of sea turtles: effects of ignoring water potential. Anthony C. Steyermark 521
Linnaeus Fund Research Reports
Reproductive cycles of male and female giant tortoises (Geochelone nigra) on the Galápagos Islands by plasma steroid analysis and ultrasound scanning. Beatrix G. Schramm, Valentine A. Lance, and Miguel Casares 523
A test of the comparative influences of constant and fluctuating incubation temperatures on phenotypes of hatchling turtles. J. Sean Doody 529
Mercury concentration in the scutes of black sea turtles, Chelonia mydas agassizii, in the Gulf of California. Stephanie M. Presti, Antonio Resendiz S. Hidalgo, Albert E. Sollod, and Jeffrey A. Seminoff 531
Turtle Poetry
Ancient Chelonians. Anders G.J. Rhodin 534
Newsnotes and Announcements
Chelonian Research Foundation Linnaeus Fund: 1999 grant recipients. Anders G.J. Rhodin 535
A fish spearing decoy in the likeness of a Blanding’s turtle (Emydoidea blandingii). Philip A. Cochran 536
A new outlet for chelonian newsnotes and announcements: Turtle and Tortoise Newsletter. Heather J. Kalb and Allen Salzberg 536
Volume 3, Number 4 – November 2000
Special Focus Issue
The Blanding’s Turtle, Emydoidea blandingii
Prologue
Editorial comment. Anders G.J. Rhodin 553
Special Focus Section
Foreword
Blanding’s Turtle Workshop summary. John J. Moriarty 555
Articles
The Blanding’s turtles (Emydoidea blandingii) of Weaver Dunes, Minnesota. Michael J. Pappas, Bruce J. Brecke, and Justin D. Congdon 557
Nesting ecology and embryo mortality: implications for hatchling success and demography of Blanding’s turtles (Emydoidea blandingii). Justin D. Congdon, Roy D. Nagle, Owen M. Kinney, Matthew Osentoski, Harold W. Avery, Richard C. van Loben Sels, and Donald W. Tinkle 569
Population structure and reproductive ecology of Blanding’s turtle (Emydoidea blandingii) in Maine, near the northeastern edge of its range. Lisa A. Joyal, Mark McCollough, and Malcolm L. Hunter, Jr. 580
Spatial ecology of Blanding’s turtle in central Minnesota. Stephen A. Piepgras and Jeffrey W. Lang 589
Occurrence, habitat selection, and movement patterns of juvenile Blanding’s turtles (Emydoidea blandingii) in Kejimkujik National Park, Nova Scotia. Natalie L. McMaster and Thomas B. Herman 602
Movement of hatchling Blanding’s turtles (Emydoidea blandingii) in Nova Scotia in response to proximity to open water: a manipulative experiment. Jennifer A. McNeil, Thomas B. Herman, and K. Lorraine Standing 611
Growth and population structure of Emydoidea blandingii from western Nebraska. David J. Germano, R. Bruce Bury, and Mary Jennings 618
Thermal ecology of Blanding’s turtle in central Minnesota. Todd D. Sajwaj and Jeffrey W. Lang 626
Results of the nest protection program for Blanding’s turtle in Kejimkujik National Park, Canada: 1987–1997. K. Lorraine Standing, Thomas B. Herman, Maria Shallow, Terry Power, and Ian P. Morrison 637
Impact of a controlled wetland drawdown on Blanding’s turtles in Minnesota. Carol Dorff Hall and Francesca J. Cuthbert 643
Restoration of wetland and upland habitat for the Blanding’s turtle, Emydoidea blandingii. Erik Kiviat, Gretchen Stevens, Robert Brauman, Sven Hoeger, Peter J. Petokas, and Garrett G. Hollands 650
Notes and Field Reports
Predation of neonate Blanding’s turtles (Emydoidea blandingii) by short-tailed shrews (Blarina brevicauda). K. Lorraine Standing, Thomas B. Herman, and Ian P. Morrison 658
Developmental abnormalities in a northeastern population of Blanding’s turtle, Emydoidea blandingii. K. Lorraine Standing, Thomas B. Herman, and Ian P. Morrison 661
Commentaries and Reviews
Commercial exploitation of Blanding’s turtle, Emydoidea blandingii, and the wood turtle, Clemmys insculpta, for the live animal trade. John P. Levell 665
Regular Contributions
Articles
A new species of long-necked turtle (Testudines: Chelidae) from the Arnhem Land Plateau, Northern Territory, Australia. Scott Thomson, Rod Kennett , and Arthur Georges 675
Reproductive ecology of the loggerhead turtle, Caretta caretta, on Fethiye and Kizilot beaches, Turkey. Oguz Türkozan 686
Do wire cages protect sea turtles from foot traffic and mammalian predators? Melissa L. Mroziak, Michael Salmon, and Kirt Rusenko 693
Habitat selection and modification by the gopher tortoise, Gopherus polyphemus, in Georgia longleaf pine forest. Melissa D. Boglioli, William K. Michener, and Craig Guyer 699
Ultrasound scanning of ovaries and eggs in Galápagos tortoises, Geochelone nigra, on Santa Cruz Island, Galápagos. Beatrix G. Schramm, Miguel Casares, and Valentine A. Lance 706
Environmental characteristics of desert tortoise (Gopherus agassizii) burrow locations in an altered industrial landscape. Jeffrey E. Lovich and Ramona Daniels 714
Local harvest of hingeback tortoises, Kinixys erosa and K. homeana, in southwestern Cameroon. Dwight P. Lawson 722
Notes and Field Reports
The green turtle, Chelonia mydas, in the Saudi Arabian Gulf. Nicolas J. Pilcher 730
Use of a marked population of diamondback terrapins (Malaclemys terrapin) to determine impacts of recreational crab pots. Margaret E. Hoyle and J. Whitfield Gibbons 735
Update on permanent residency, persistence, and longevity in a 35-year study of a population of three-toed box turtles in Missouri. Elizabeth R. Schwartz 737
A translocation strategy for confiscated pancake tortoises. Luke James Harmon 738
Aquatic home ranges of female western pond turtles, Clemmys marmorata, at two sites in southern California. Robert H. Goodman, Jr. and Glenn R. Stewart 743
The identification of the holotype of Chelodina oblonga (Testudines: Chelidae) with a discussion of taxonomic implications. Scott Thomson 745
Observations of male green turtles (Chelonia mydas) on the nesting beach at Tortuguero National Park, Costa Rica. Sebastian Troëng 749
Predation of green (Chelonia mydas) and leatherback (Dermochelys coriacea) turtles by jaguars (Panthera onca) at Tortuguero National Park, Costa Rica. Sebastian Troëng 751
Mud accumulation in nesting aquatic turtles (Emydidae) in Illinois. John K. Tucker and Daniel A. Warner 753
Commentaries and Reviews
Sexing juvenile sea turtles: is there an accurate and practical method? Thane Wibbels, David Wm. Owens, and Colin J. Limpus 756
A response to Nicholas Mrosovsky’s Sustainable Use of Hawksbill Turtles: Contemporary Issues in Conservation. Peter C.H. Pritchard 761
Conservation and cultural diversity: reply to Pritchard. N. Mrosovsky 767
Turtle Poetry
Tragedy of the road. Don Blanding 770
Volume 4 (2001–2005)
Volume 4, Number 1 – September 2001
Prologue
Editorial comment. Anders G.J. Rhodin 3
Articles
Observations on body size, sympatry, and niche divergence in softshell turtles (Trionychidae). Peter C.H. Pritchard 5
Sexing young free-ranging desert tortoises (Gopherus agassizii) using external morphology. James L. Boone and Eric A. Holt 28
The green turtle, Chelonia mydas, in Queensland, Australia: the Bramble Cay rookery in the 1979–1980 breeding season. Colin J. Limpus, David Carter, and Mark Hamann 34
Genetic population structure of two threatened South American river turtle species, Podocnemis expansa and Podocnemis unifilis. Brian C. Bock, Vivian P. Páez, and Matthew M. White 47
Morphometric model for sex assessment in hatchling olive ridley sea turtles. Jesús Emilio Michel-Morfin, Victor M. Gómez Muñoz, and Carmen Navarro Rodríguez 53
Deforestation and habitat loss for the Malagasy flat-tailed tortoise from 1963 through 1993. Stephen T. Tidd, John E. Pinder III, and Gary W. Ferguson 59
Demographic and ecological factors affecting conservation and management of the diamondback terrapin (Malaclemys terrapin) in South Carolina. J. Whitfield Gibbons, Jeffrey E. Lovich, Anton D. Tucker, Nancy N. FitzSimmons, and Judith L. Greene 66
Seasonal movements and reproduction in the striped mud turtle (Kinosternon baurii) from the southern Everglades, Florida. Walter E. Meshaka, Jr. and Eric Blind 75
Use of thermal springs for aquatic basking by the pig-nosed turtle, Carettochelys insculpta. J. Sean Doody, Rachel A. Sims, and Arthur Georges 81
A system-of-equations growth function for southern Great Barrier Reef green sea turtles. Milani Chaloupka 88
Some ecological parameters of the wood turtle, Clemmys insculpta, in southeastern Pennsylvania. Carl H. Ernst 94
Sexual dimorphism and morphological differentiation in European pond turtle (Emys orbicularis) populations from northwestern Spain. César Ayres Fernández and Adolfo Cordero Rivera 100
Migration routes and destination characteristics of post-nesting hawksbill turtles satellite-tracked from Barbados, West Indies. Julia A. Horrocks, Lotus A. Vermeer, Barry Krueger, Michael Coyne, Barbara A. Schroeder, and George H. Balazs 107
Terrestrial habitat use by aquatic turtles from a seasonally fluctuating wetland: implications for wetland conservation boundaries. Kurt A. Buhlmann and J. Whitfield Gibbons 115
Trace metal concentrations in blood of the Kemp’s ridley sea turtle (Lepidochelys kempii). Lisa O. Kenyon, André M. Landry, Jr., and Gary A. Gill 128
Habitat use and movements of radiotagged Blanding’s turtles (Emydoidea blandingii) in a suburban landscape. Cory S. Rubin, Richard E. Warner, and Daniel R. Ludwig 136
The loggerhead turtle, Caretta caretta, in Queensland: breeding migrations and fidelity to a warm temperate feeding area. Colin J. Limpus and Duncan J. Limpus 142
Notes and Field Reports
Notes on the distribution, life history, and exploitation of turtles in Sulawesi, Indonesia, with emphasis on Indotestudo forstenii and Leucocephalon yuwonoi. Steven G. Platt, Robert J. Lee, and Michael W. Klemens 154
Kyphosis in a musk turtle (Sternotherus odoratus) from Ontario, Canada. Raymond A. Saumure 159
Surface ultrastructure and retention of the egg caruncle in the European pond turtle, Emys orbicularis. Slawomir Mitrus 160
Response of juvenile softshell turtles (Apalone mutica) in a thermal gradient. Alan V. Nebeker and R. Bruce Bury 162
Husbandry, behavior, and captive breeding of the Nama padloper, Homopus bergeri, from southwestern Namibia. Alfred Schleicher and Victor J.T. Loehr 165
Mycoflora identified from failed green (Chelonia mydas) and loggerhead (Caretta caretta) sea turtle eggs at Heron Island, Australia. Andrea D. Phillott, C. John Parmenter, and Colin J. Limpus 170
A tortoise survey of Shwe Settaw Wildlife Sanctuary, Myanmar, with notes on the ecology of Geochelone platynota and Indotestudo elongata. Steven G. Platt, Saw Tun Khaing, Win Ko Ko, and Kalyar 172
Summertime foraging and feeding by immature loggerhead sea turtles (Caretta caretta) from Georgia. Michael G. Frick, Kristina L. Williams, and Lianne Pierrard 178
Winter survey of a gopher tortoise population in South Carolina. Tracey D. Tuberville and Michael E. Dorcas 182
Penis displays of snapping turtles (Chelydra serpentina) in response to handling: defensive or displacement behavior? Shane R. de Solla, Melinda Portelli, Holly Spiro, and Ronald J. Brooks 187
Maximum size and mass of the ploughshare tortoise, Geochelone yniphora. Miguel Pedrono and Tim Markwell 190
Nesting biology of the sea turtles of St. Kitts, West Indies. Joseph A. Butler 191
Assessment of sea turtle mortality rates in the Bahía Magdalena region, Baja California Sur, México. Susan C. Gardner and Wallace J. Nichols 197
Diet of the softshell turtle, Trionyx triunguis, in an Afrotropical forested region. Godfrey C. Akani, Dario Capizzi, and Luca Luiselli 200
Feeding of Dermochelys coriacea on medusae in the northwest Atlantic. Michael C. James and Thomas B. Herman 202
Over three decades of persistence of a small and apparently isolated population of painted turtles (Chrysemys picta) in a Kentucky reservoir. Peter V. Lindeman and A. Floyd Scott 206
On the possible occurrence of the marginated tortoise, Testudo marginata, in Turkey. Oguz Türkozan, Yusuf Kumlutas, and Çetin Ilgaz 208
Commentaries and Reviews
An overview of the North American turtle genus Clemmys Ritgen, 1828. Carl H. Ernst 211
Relative importance of thermal and nonthermal factors on the incubation period of sea turtle eggs. Matthew H. Godfrey and N. Mrosovsky 217
Linnaeus Fund Research Reports
Age estimation of Eretmochelys imbricata by schlerochronology of carapacial scutes. Anton D. Tucker, Damien Broderick, and Lorelei Kampe 219
Investigations of the ecology of Graptemys versa and Pseudemys texana in southcentral Texas. Peter V. Lindeman 223
Molecular systematics of emydine turtles. Chris R. Feldman and James Ford Parham 224
Genetic relatedness of populations of the mud turtle, Kinosternon baurii, from mainland Florida and the upper and Lower Florida Keys. Dawn S. Wilson and Stephen A. Karl 228
Turtle Poetry
Playa Grande beach. Michael H.J. Rhodin 231
Volume 4, Number 2 – December 2002
Special Focus Issue
The Gopherine Tortoises, Gopherus spp.
Prologue
Editorial comment. Anders G.J. Rhodin 247
Articles
The Desert Tortoise, Gopherus agassizii
Defining the desert tortoise(s): our first priority for a coherent conservation strategy. Kristin H. Berry, David J. Morafka, and Robert W. Murphy 249
Is Gopherus agassizii a desert-adapted tortoise, or an exaptive opportunist? Implications for tortoise conservation. David J. Morafka and Kristin H. Berry 263
Reproduction in a desert tortoise (Gopherus agassizii) population on the Beaver Dam Slope, Washington County, Utah. Ann M. McLuckie and Rick A. Fridell 288
Reproduction of Gopherus agassizii in the Sonoran Desert, Arizona. Roy C. Averill-Murray 295
The annual reproductive cycle of the male and female desert tortoise: physiology and endocrinology. Valentine A. Lance and David C. Rostal 302
Chronology of sex determination in the desert tortoise (Gopherus agassizii). David C. Rostal, Thane Wibbels, Janice S. Grumbles, Valentine A. Lance, and James R. Spotila 313
Energy and water balance, diet, and reproduction of female desert tortoises (Gopherus agassizii). Brian Thomas Henen 319
Alien plants and fire in desert tortoise (Gopherus agassizii) habitat of the Mojave and Colorado deserts. Matthew L. Brooks and Todd C. Esque 330
Selective spring foraging by juvenile desert tortoises (Gopherus agassizii) in the Mojave Desert: evidence of an adaptive nutritional strategy. Olav T. Oftedal, Scott Hillard, and David J. Morafka 341
Diet selection by the desert tortoise in relation to the flowering phenology of ephemeral plants. W. Bryan Jennings 353
An experimental assessment of color, calcium, and insect dietary preferences of captive juvenile desert tortoises (Gopherus agassizii). Chris L. Okamoto 359
A landscape sampling protocol for estimating distribution and density patterns of desert tortoises at multiple spatial scales. Anthony J. Krzysik 366
Regional desert tortoise monitoring in the Upper Virgin River Recovery Unit, Washington County, Utah. Ann M. McLuckie, Deborah L. Harstad, Jesse W. Marr, and Rick A. Fridell 380
Spatial organization of desert tortoises and their burrows at a landscape scale. Jeffrey J. Duda, Anthony J. Krzysik, and Joel M. Meloche 387
Patterns of burrow use by desert tortoises (Gopherus agassizii) in southcentral Nevada. Kurt R. Rautenstrauch, Danny L. Rakestraw, Greg A. Brown, James L. Boone, and Patrick E. Lederle 398
Comparative dispersion of neonate and headstarted juvenile desert tortoises (Gopherus agassizii): a preliminary assessment of age effects. Lisa C. Hazard and David J. Morafka 406
Shell kinesis in juvenile desert tortoises, Gopherus agassizii. Marsha L. Marolda 410
Using growth ring counts to age juvenile desert tortoises (Gopherus agassizii) in the wild. Kristin H. Berry 416
A condition index for the desert tortoise (Gopherus agassizii). Kenneth A. Nagy, Brian T. Henen, Devesh B. Vyas, and Ian R. Wallis 425
Effects on survival of desert tortoises (Gopherus agassizii) urinating during handling. Roy C. Averill-Murray 430
Deaths of desert tortoises following periods of drought and research manipulation. Kristin H. Berry, E. Karen Spangenberg, Bruce L. Homer, and Elliott R. Jacobson 436
Impacts of vehicle road traffic on desert tortoise populations with consideration of conservation of tortoise habitat in southern Nevada. Karin von Seckendorff Hoff and Ronald William Marlow 449
Comparison of desert tortoise (Gopherus agassizii) populations in an unused and off-road vehicle area in the Mojave Desert. R. Bruce Bury and Roger A. Luckenbach 457
The Gopher Tortoise, Gopherus polyphemus
Reserve area requirements for gopher tortoises (Gopherus polyphemus). Jeannine Ott Eubanks, Jeff W. Hollister, Craig Guyer, and William K. Michener 464
A comparison of GIS and survey estimates of gopher tortoise habitat and numbers of individuals in Florida. Earl D. McCoy, Beth Stys, and Henry R. Mushinsky 472
Population biology of the gopher tortoise (Gopherus polyphemus) in southeast Georgia. David C. Rostal and Douglas N. Jones, Jr. 479
The Texas Tortoise, Gopherus berlandieri
Home range and dispersal of Texas tortoises, Gopherus berlandieri, in a managed thornscrub ecosystem. Richard T. Kazmaier, Eric C. Hellgren, and Donald C. Ruthven, III 488
Mycoplasmal Disease in Gopherus spp.
Application of diagnostic tests for mycoplasmal infections of desert and gopher tortoises, with management recommendations. Daniel R. Brown, Isabella M. Schumacher, Grace S. McLaughlin, Lori D. Wendland, Mary B. Brown, Paul A. Klein, and Elliott R. Jacobson 497
Turtle Poetry
Population density studies. Marian L. Griffey 508
Volume 4, Number 3 – December 2003
Prologue
Editorial comment. Anders G.J. Rhodin 521
Articles
Reproductive biology and conservation status of the loggerhead sea turtle (Caretta caretta) in Espírito Santo State, Brazil. Cecília Baptistotte, João C.A. Thomé, and Karen A. Bjorndal 523
Post-nesting movements and behavior of loggerhead sea turtles (Caretta caretta) departing from east-central Florida nesting beaches. C. Kenneth Dodd, Jr. and Richard Byles 530
Previously unrecognized original type specimens of American turtles collected by John Le Conte in 1828. Roger Bour 537
Emergence success of flatback sea turtles (Natator depressus) at Fog Bay, Northern Territory, Australia. Sean J. Blamires and Michael L. Guinea 548
Influence of nest site selection on predation of flatback sea turtle (Natator depressus) eggs by varanid lizards in northern Australia. Sean J. Blamires, Michael L. Guinea, and Robert I.T. Prince 557
Diagnostic characteristics in lower Tennessee River populations of the map turtles Graptemys pseudogeographica and Graptemys ouachitensis. Peter V. Lindeman 564
Activity and thermal ecology of the box turtle, Terrapene ornata, at its southwestern range limit in Arizona. Michael V. Plummer 569
Daily and seasonal activity patterns and movements of juvenile box turtles (Terrapene carolina bauri) on Egmont Key, Florida. Alison Hamilton Jennings 578
Nesting and conservation of the hawksbill turtle, Eretmochelys imbricata, in the Pearl Cays, Nicaragua. Cynthia J. Lagueux, Cathi L. Campbell, and William A. McCoy 588
Comparative study of the nesting behavior of the green turtle, Chelonia mydas, during high- and low-density nesting periods at Ras Al-Hadd Reserve, Oman. A.Y.A. AlKindi, I.Y. Mahmoud, H.M. Al-Gheilani, C.S. Bakheit, A.A. Al-Habsi, and A. Al-Kiyumi 603
Status of the flatback sea turtle (Natator depressus) rookery on Crab Island, Australia, with notes on predation by crocodiles. Ronald W. Sutherland and Elizabeth G. Sutherland 612
Marine turtle conservation on Holbox Island, Quintana Roo, Mexico: the fishermen’s perceptions. Sonja Macys and George N. Wallace 620
Influence of geological factors on reproductive aspects of Podocnemis unifilis (Testudines, Pelomedusidae) on the Javaés River, Araguaia National Park, Brazil. Paulo Dias Ferreira Júnior, Adriana Malvasio, and Odimar Santos Guimarães 626
Morphological variation in the endangered Alabama red-bellied cooter (Pseudemys alabamensis) and taxonomic status of a population in Mississippi. Christopher J. Leary, James L. Dobie, Thomas M. Mann, and Peter S. Floyd 635
Biological prioritization of Asian countries for turtle conservation. Bryan L. Stuart and John Thorbjarnarson 642
Status of Testudo kleinmanni and T. werneri in Egypt. Sherif M. Baha El Din, Omar Attum, and Mindy Baha El Din 648
Home range and seasonal movements of the wood turtle (Glyptemys insculpta) in southern New Hampshire. Sheila E. Tuttle and David M. Carroll 656
Reproductive ecology of Podocnemis erythrocephala (Testudines: Podocnemididae) in the lower Inírida River, Colombia. Olga Victoria Castaño-Mora, Pedro Alberto Galvis-Peñuela, and Juan Gabriel Molano 664
Notes and Field Reports
Akinesis and plastral scute homologies in Sternotherus (Testudines: Kinosternidae). Peter C.H. Pritchard 671
Marine turtle nesting activity at Playa Naranjo, Santa Rosa National Park, Costa Rica, for the 1998–1999 season. Dana L. Drake, Jocelyn E. Behm, Meghan A. Hagerty, Philippe A. Mayor, Seth J. Goldenberg, and James R. Spotila 675
Ecology and conservation status of the Arakan forest turtle, Heosemys depressa, in western Myanmar. Steven G. Platt, Win Ko Ko, Kalyar, Myo Myo, Lay Lay Khaing, and Thomas R. Rainwater 678
Comparative ultrastructural carapace morphology in three freshwater turtles. David A. Wooten 682
Diet of sea turtles in southern Brazil. Leandro Bugoni, Lígia Krause, and Maria Virgínia Petry 685
Radiotransmitter attachment technique for box turtles (Terrapene spp.). Karen A. Wilson, Paul M. Cavanagh, and Jeffrey Villepique 688
Rediscovery of the critically endangered river terrapin, Batagur baska, in Cambodia, with notes on occurrence, reproduction, and conservation status. Steven G. Platt, Bryan L. Stuart, Heng Sovannara, Long Kheng, Kalyar, and Heng Kimchhay 691
Effects of sand hardness and human beach use on emergence success of loggerhead sea turtles on Yakushima Island, Japan. Hiromi Kudo, Akinobu Murakami, and Satomi Watanabe 695
Reproductive cycle of male snapping turtles (Chelydra serpentina) in southeastern Virginia. Lauren L. Glesenkamp, George R. Zug, and Joseph C. Mitchell 697
Natural hybridization between a Kemp’s ridley (Lepidochelys kempii) and loggerhead sea turtle (Caretta caretta) confirmed by molecular analysis. Robert C. Barber, Clark T. Fontaine, Joseph P. Flanagan, and Edward E. Louis, Jr. 701
Double-chambered nest cavities in the leatherback turtle. Stephanie Kamel and N. Mrosovsky 705
Summer movements and home range of the cooter turtle, Pseudemys concinna, in Illinois. Michael J. Dreslik, Andrew R. Kuhns, Christopher A. Phillips, and Benjamin C. Jellen 706
Comparative efficiency of different sampling techniques to obtain DNA from freshwater turtles. Nathalie Tessier and François-Joseph Lapointe 710
Effect of translocation on egg viability of the giant Amazon river turtle, Podocnemis expansa. Kelly Bonach, Maristela P. Miranda-Vilela, Marcelo C. Alves, and Luciano M. Verdade 712
First record of a gravid marine turtle from Chile. Angélica González, Leyla Miranda, and Juan Carlos Ortiz 716
Repetitive data-logger attachments to sea turtles using a new quick-release method. Sandra Storch and Solvin Zankl 717
Preliminary observations on reproductive parameters of the Sulawesi forest turtle (Leucocephalon yuwonoi) in captivity. Charles J. Innis 720
Population ecology of musk turtles (Sternotherus odoratus) in a lake in Virginia. Alison M. Holinka, Radhika Dave, and Randolph M. Chambers 721
Axial bifurcation in a bicephalic Chelonia mydas embryo. Cheong H. Diong, Lenny K.A. Tan, and Charles M.U. Leh 725
Commentaries and Reviews
Turtle management as scientific experimentation. William E. Magnusson 728
Linnaeus Fund Research Reports
Distribution, status, and biology of the radiated tortoise, Geochelone radiata, in southwest Madagascar. Thomas E.J. Leuteritz and Rollande Ravolanaivo 730
Domestic trade of tortoises and freshwater turtles in Cambodia. Rohan H.P. Holloway 733
Assessing the impact of human hunting activities on populations of forest tortoises (genus Kinixys) in the Niger Delta, Nigeria. Luca Luiselli 735
Interpond movement patterns of Chrysemys picta picta (eastern painted turtles) in Virginia. David R. Bowne 739
Turtle Poetry
The egg and the machine. Robert Frost 742
Volume 4, Number 4 – April 2005
Special Focus Issue
The Kemp’s Ridley Sea Turtle, Lepidochelys kempii
Prologue
Editorial comment. Anders G.J. Rhodin 757
Special Focus Section
Foreword
Introduction to the Kemp’s ridley focus issue. Peter C.H. Pritchard and David W. Owens 759
Articles
Status of the Kemp’s ridley sea turtle, Lepidochelys kempii. René Márquez-M., Patrick M. Burchfield, Juan Díaz-F., Manuel Sánchez-P., Miguel Carrasco-A., Carmen Jiménez-Q., Alma Leo-P., Rafael Bravo-G., and Jaime Peña-V. 761
A population model to estimate recovery time, population size, and management impacts on Kemp’s ridley sea turtles. Selina S. Heppell, Deborah T. Crouse, Larry B. Crowder, Sheryan P. Epperly, Wendy Gabriel, Terry Henwood, René Márquez, and Nancy B. Thompson 767
Time-series analyses of the relationship between nesting frequency of the Kemp’s ridley sea turtle and meteorological conditions. María del Carmen Jiménez-Quiroz, Anatoly Filonov, Irina Tereshchenko, and René Márquez-Millán 774
Reproductive parameters of the Kemp’s ridley sea turtle (Lepidochelys kempii) at Rancho Nuevo, Tamaulipas, Mexico. Wayne N. Witzell, Alfredo Salgado-Quintero, and Manuel Garduño-Dionte 781
Seasonal reproductive biology of the Kemp’s ridley sea turtle (Lepidochelys kempii): comparison of captive and wild populations. David C. Rostal 788
Population characteristics of Kemp’s ridley sea turtles in nearshore waters of the upper Texas and Louisiana coasts. André M. Landry, Jr., David T. Costa, F. Leonard Kenyon, II, and Michael S. Coyne 801
Kemp’s ridley sea turtle movements and migrations. Maurice L. Renaud and Jo A. Williams 808
Movements and home ranges of adult male Kemp’s ridley sea turtles (Lepidochelys kempii) in the Gulf of Mexico investigated by satellite telemetry. Donna J. Shaver, Barbara A. Schroeder, Richard A. Byles, Patrick M. Burchfield, Jaime Peña, René Márquez, and Hector J. Martinez 817
Developmental biology and ecology of the Kemp’s ridley sea turtle, Lepidochelys kempii, in the eastern Gulf of Mexico. Jeffrey R. Schmid and William J. Barichivich 828
Texans, turtles, and the early Kemp’s ridley population restoration project, 1963–67. Patrick M. Burchfield 835
Head-starting the Kemp’s ridley sea turtle, Lepidochelys kempii, at the NMFS Galveston Laboratory, 1978–1992: a review. Clark Fontaine and Donna Shaver 838
Analysis of the Kemp’s ridley imprinting and headstart project at Padre Island National Seashore, Texas, 1978–88, with subsequent nesting and stranding records on the Texas coast. Donna J. Shaver 846
On the sex ratio of juvenile Lepidochelys kempii in Georgia. Carol Ruckdeschel, C. Robert Shoop, and Robert D. Kenney 860
Diet analysis of Kemp’s ridley sea turtles (Lepidochelys kempii) in Virginia. Erin E. Seney and John A. Musick 864
Western North Atlantic waters: crucial developmental habitat for Kemp’s ridley and loggerhead sea turtles. Stephen J. Morreale and Edward A. Standora 872
Climatic and oceanographic factors affecting daily patterns of juvenile sea turtle cold-stunning in Cape Cod Bay, Massachusetts. Brett M. Still, Curtice R. Griffin, and Robert Prescott 883
Regular Contributions
Articles
Assessment of the threatened status of Testudo kleinmanni Lortet, 1883 (Testudines: Testudinidae) for the IUCN Red List. Jarmo Perälä 891
Conserving sea turtles in Canada: successful community-based collaboration between fishers and scientists. Kathleen Martin and Michael C. James 899
Ecology of the Jamaican slider turtle (Trachemys terrapen), with implications for conservation and management. Tracey D. Tuberville, Kurt A. Buhlmann, Rhema Kerr Bjorkland, and Doug Booher 908
Nesting activity and reproductive output of loggerhead sea turtles, Caretta caretta, over 19 seasons (1984–2002) at Laganas Bay, Zakynthos, Greece: the largest rookery in the Mediterranean. Dimitris Margaritoulis 916
Documenting the value of volunteer effort for sea turtle conservation in South Carolina. Sally R. Hopkins-Murphy and Joan S. Seithel 930
Notes and Field Reports
Bifid tail regeneration in a turtle, Emydura sp. (Testudines: Chelidae). Gerald Kuchling 935
Thermoregulation in captive Indian Ocean giant tortoises. Justin Gerlach 937
Noteworthy records and exploitation of chelonians from the Ayeyarwady, Chindwin, and Dokhtawady rivers, Myanmar. Steven G. Platt, Win Ko Ko, Lay Lay Khaing, Khin Myo Myo, Tint Lwin, Thanda Swe, Kalyar, and Thomas R. Rainwater 942
Distribution, natural history, and exploitation of Leucocephalon yuwonoi in central Sulawesi, Indonesia. Cris Hagen and Iwalani Y.S. Ching 948
Genetic analysis of mitochondrial DNA variation in eastern and western African spurred tortoises, Geochelone sulcata. Barbara Livoreil and Antoinette C. van der Kuyl 951
Distribution of the black-breasted leaf turtle (Geoemyda spengleri) on Hainan Island, China. Shi Haitao 954
Occurrence of diamondback terrapins, Malaclemys terrapin, on Bermuda: native or introduced? John Davenport, Anne F. Glasspool, and Lisa Kitson 956
Turtle Poetry
Turtles. Author Unknown 960
Available at www.chelonianjournals.org
