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Chelonian Conservation and Biology
International Journal of Turtle and Tortoise Research
(ISSN 1071-8443)

Volumes 1–4 (1993–2005)


Volume 1 (1993–1995)


Volume 1, Number 1 – November 1993


Editorial: A new turtle and tortoise journal. JOHN L. BEHLER, PETER C.H. PRITCHARD, AND ANDERS G.J. RHODIN     2


Conservation biology of the pig-nosed turtle, Carettochelys insculpta. ARTHUR GEORGES AND MARK ROSE     3

Status and exploitation of the Madagascan big-headed turtle, Erymnochelys madagascariensis. GERALD KUCHLING AND RUSSELL A. MITTERMEIER     13

Distribution, osteology, and natural history of the Asian giant softshell turtle, Pelochelys bibroni, in Papua New Guinea. ANDERS G.J. RHODIN, RUSSELL A. MITTERMEIER, AND PHILIP M. HALL     19

Carapacial pankinesis in the Malayan softshell turtle, Dogania subplana. PETER C.H. PRITCHARD     31

On growth, sexual dimorphism, and the general ecology of the African spurred tortoise, Geochelone sulcata, in Mali. MICHAEL R.K. LAMBERT     37

Notes and Field Reports

Range extension for Emydura subglobosa in Papua New Guinea. ANDERS G.J. RHODIN     47

A ranching project for freshwater turtles in Costa Rica. PETER C.H. PRITCHARD     48

New localities for Pyxis planicauda in west-central Madagascar. JOHN L. BEHLER, QUENTIN M.C. BLOXAM, EDMOND R. RAKOTOVAO, AND HERILALA J.A.R. RANDRIAMAHAZO     49

Reunion Island – still a land of tortoises. JUAN GONZALEZ GONZALEZ     51


On the distribution, relative abundance and protection of tortoises in Bulgaria. VLADIMIR A. BESHKOV [1984]. Translation from Bulgarian arranged by JAMES R. BUSKIRK     53

Newsnotes and Announcements

International conference on conservation, restoration, and management of turtles and tortoises. MICHAEL W. KLEMENS, KRISTIN H. BERRY, AND SUZANNE DOHM     63

Bibliographic database of chelonian literature established. ANDERS G.J. RHODIN, KENT R. BEAMAN, AND LORI A. DICONZA     64

Chelonian Research Foundation Linnaeus Fund: 1992 grant recipients. ANDERS G.J. RHODIN     64


IUCN/SSC Tortoise and Freshwater Turtle Specialist Group 1993 membership list.     67


Volume 1, Number 2 – July 1994




The Mary River tortoise: a new genus and species of short-necked chelid from Queensland, Australia (Testudines; Pleurodira). JOHN CANN AND JOHN M. LEGLER     81

Activity patterns and habitat use of box turtles (Terrapene carolina bauri) on a Florida island, with recommendations for management. C. KENNETH DODD, JR., RICHARD FRANZ, AND LORA L. SMITH     97

The ecology of sea beach nesting in slider turtles (Trachemys scripta venusta) from Caribbean Costa Rica. DON MOLL     107

Morphometric analysis and taxonomy of cooter and red-bellied turtles in the North American genus Pseudemys (Emydidae). MICHAEL E. SEIDEL     117

Comments on Chinemys palaeannamitica and certain other chelonian taxa based upon material from archaeological sites. PETER C.H. PRITCHARD     131

Notes and Field Reports

Husbandry and captive breeding of the parrot-beaked tortoise, Homopus areolatus. JAMES E. BARZYK     138

Thermal limits of incubation in embryos of softshell turtles (Apalone mutica). MICHAEL V. PLUMMER, CHRIS E. SHADRIX, AND REBEKAH C. COX     141

Reproduction of the cabeçudo, Peltocephalus dumerilianus, in the Biological Reserve of Rio Trombetas, Pará, Brazil. RICHARD C. VOGT, VITOR HUGO CANTARELLI, AND ALBERTO GUERREIRO DE CARVALHO     145

Maximum size and clutch size records for eastern painted turtles, Chrysemys picta picta, from mid-coastal Maine. ANDERS G.J. RHODlN AND GLEN H. MITTELHAUSER     148


Status and conservation prospects of Testudo graeca L. inhabiting the Black Sea coast of the Caucasus. ALEXANDER A. INOZEMTSEV AND SOLOMON L. PERESHKOLNlK. [1985]. Translation from Russian by MARINA PROUTKINA, arranged by DEWEY I. WALLACE     151

Commentaries and Reviews

Temperature controlled sex determination as a tool for turtle conservation. RICHARD C. VOGT     159

Newsnotes and Announcements

Turtle and tortoise papers presented at the Second World Congress of Herpetology.     163

Chelonia Neotropica – a new turtle and tortoise newsletter from Brazil. ALINE TRISTAO BERNARDES    165

Chelonian Research Foundation Linnaeus Fund: 1993 grant recipients. ANDERS G.J. RHODIN     166


Volume 1, Number 3 – January 1995




Movements and home range of hatchling and yearling gopher tortoises, Gopherus polyphemus. JOSEPH A. BUTLER, RAY D. BOWMAN, TODD W. HULL, AND SCOTT SOWELL     173

Reproductive biology of the chopontil, Claudius angustatus (Testudines: Kinosternidae), in southern Veracruz, Mexico. OSCAR A. FLORES-VILLELA AND GEORGE R. ZUG     181

Early post-emergent behavior and habitat selection in hatchling Blanding’s turtles, Emydoidea blandingii, in Massachusetts. BRIAN O. BUTLER AND TERRY E. GRAHAM     187

An assessment of the effects of commercial exploitation on the pancake tortoise, Malacochersus tornieri, in Tanzania. MICHAEL W. KLEMENS AND DON MOLL     197

Habitat use and population parameters of the spotted turtle, Clemmys guttata, a Species of Special Concern in Massachusetts. TERRY E. GRAHAM     207

Observations and morphometric data on the Namaqualand speckled tortoise, Homopus signatus signatus (Gmelin, 1789), in South Africa. NICOLAS BAYOFF     215

Notes and Field Reports

Exploitation of the alligator snapping turtle, Macroclemys temminckii, in Louisiana: a case study. KEVIN SLOAN AND JEFFREY E. LOVICH     221

Turtles at a market in western Yunnan: possible range extensions for some southern Asiatic chelonians in China and Myanmar. GERALD KUCHLING     223

Reproductive biology of the Indian roofed turtle, Kachuga tecta, in Bangladesh. MD. LOKMAN HOSSAIN AND MD. SOHRAB UDDIN SARKER     226

Incubation period and sex ratio of Hermann’s tortoise, Testudo hermanni boettgeri. BERT T. EENDEBAK     227

Growth of head-started Kemp’s ridley sea turtles (Lepidochelys kempii) following release. CHARLES W. CAILLOUET, JR., CLARK T. FONTAINE, SHARON A. MANZELLA-TIRPAK, AND THEODORE D. WILLIAMS     231


Population density of the steppe tortoise in some regions of the Almaty and Taldyqorghan Districts, Kazakhstan. RUDOLF A. KUBYKIN. [1988]. Translation from Russian by CHERYL A. WHlTE, arranged by JAMES R. BUSKIRK     235

Commentaries and Reviews

Manipulating sex ratios: turtle speed ahead! N. MROSOVSKY AND MATTHEW H. GODFREY     238

Workshop on the ecology, status, and management of the diamondback terrapin (Malaclemys terrapin), Savannah River Ecology Laboratory, 2 August 1994: final results and recommendations. Compiled by RICHARD A. SEIGEL AND J. WHITFIELD GIBBONS     240

Turtle issues at CITES 1994. PETER C.H. PRITCHARD     243

In Memoriam. Leo Brongersma — an appreciation. PETER C.H. PRITCHARD     244

Newsnotes and Announcements

Chelonian Research Foundation Linnaeus Fund: 1994 grant recipients. ANDERS G.J. RHODIN     248

Research activities of the Sea Turtle Research Unit (SEATRU) of Universiti Pertanian Malaysia. SEA TURTLE RESEARCH UNIT     248

Centre de Reproducció de Tortugues de l’Albera in Spain. RAMON MASCORT BRUGAROLAS     249

New publication on turtle and tortoise trade in the United States. ALLEN SALZBERG     250

Chelonian Research Monographs, a new publication series by Chelonian Research Foundation. ANDERS G.J. RHODIN     250


Volume 1, Number 4 – August 1995


On geographical size variation, growth, and sexual dimorphism of the leopard tortoise, Geochelone pardalis, in Somaliland. MICHAEL R.K. LAMBERT     269

Mycoplasmosis and the desert tortoise (Gopherus agassizii) in Las Vegas Valley, Nevada. ELLIOTT R. JACOBSON, MARY B. BROWN, ISABELLA M. SCHUMACHER, BOBBY R. COLLINS, RICHARD K. HARRIS, AND PAUL A. KLEIN     279

Survival of head-started Kemp’s ridley sea turtles (Lepidochelys kempii) released into the Gulf of Mexico or adjacent bays. CHARLES W. CAILLOUET, JR., CLARK T. FONTAINE, SHARON A. MANZELLA-TIRPAK, AND DONNA J. SHAVER     285

Geographic variation in the matamata turtle, Chelus fimbriatus, with observations on its shell morphology and morphometry. MARCELO R. SANCHEZ-VILLAGRA, PETER C.H. PRITCHARD, ALFRED PAOLILLO, AND OMAR J. LINARES     293

Redescription and neotype designation of Pelochelys bibroni from southern New Guinea (Testudines: Trionychidae). ROBERT G. WEBB     301

A new batagurid turtle from northern Sulawesi, Indonesia. WILLIAM P. MCCORD, JOHN B. IVERSON, AND BOEADI     311

Notes and Field Reports

Salvage of eggs from road-killed red-eared sliders, Trachemys scripta elegans. JOHN K. TUCKER     317

Repatriation of radiated tortoises, Geochelone radiata, from Reunion Island to Madagascar. STEPHANE BOULLAY     319

Observations on feeding habits of Hydromedusa maximiliani (Testudines: Chelidae) in southeastern Brazil. FRANCO LEANDRO SOUZA AND AUGUSTO SHINYA ABE     320

The date of publication of Gray’s Catalogue of Shield Reptiles. ROBERT G. WEBB     322


Results of studies on the chelonians of Albania. IDRIZ HAXHIU. [1985]. Translation from Albanian by JAMES R. BUSKIRK     324

Current data on the chelonians of Albania. IDRIZ HAXHIU     326

Commentaries and Reviews

A clarification on the activities of Projeto TAMAR, Brazil. CECILIA BAPTISTOTTE     328

Systematics of the Pseudemys concinna-floridana complex (Testudines: Emydidae): an alternative interpretation. DALE R. JACKSON     329

How many species of cooter turtles and where is the scientific evidence? – a reply to Jackson. MICHAEL E. SEIDEL     333

Book Review: ACUNA MESEN, R.A. 1993. Las Tortugas Continentales de Costa Rica. San Jose, Costa Rica: Editorial ICER, 58 pp. MAHMOOD SASA     336

Newsnotes and Announcements

Turtle and tortoise papers presented at the International Congress of Chelonian Conservation.     338


Volume 2 (1996–1997)


Volume 2, Number 1 – February 1996




Overwintering in the nest chamber by hatchling painted turtles, Chrysemys picta, in northern New Jersey. JOSEPH A. DEPARI     5

Reproductive biology of a flood-season nesting freshwater turtle of the northern neotropics: Dermatemys mawii in Belize. JOHN POLISAR     13

Ecological characteristics of the pancake tortoise, Malacochersus tornieri, in Tanzania. DON MOLL AND MICHAEL W. KLEMENS     26

Observations on ecological changes threatening a population of Testudo graeca graeca in the Souss Valley, southern Morocco. JANE R. BAYLEY AND ANDY C. HIGHFIELD     36

Seasonal variation of embryonic diapause in the striped mud turtle (Kinosternon baurii) and general considerations for conservation planning. MICHAEL A. EWERT AND DAWN S. WILSON     43

A six season study of marine turtle nesting at Praia do Forte, Bahia, Brazil, with implications for conservation and management. MARIA ANGELA MARCOVALDI AND ANTONIO LAURENT     55

A survey of organized turtle watch participants on sea-turtle nesting beaches in Florida. STEVE A. JOHNSON, KAREN A. BJORNDAL, AND ALAN B. BOLTEN     60

Observations on the natural history and exploitation of the turtles of Nepal, with life history notes on Melanochelys trijuga. JOSEPH C. MITCHELL AND ANDERS G.J. RHODIN     66

Notes and Field Reports

The origin of marine turtles: a pluralistic view of evolution. JOSEPH J. KINNEARY     73

Occurrence and diet of juvenile loggerhead sea turtles, Caretta caretta, in the northwestern Gulf of Mexico. PAMELA T. PLOTKIN     78

First record of Heosemys spinosa from the Philippines, with biogeographic notes. INDRANEIL DAS     80

Neural bones in Australian chelid turtles. SCOTT THOMSON AND ARTHUR GEORGES     82

Temperature-dependent sex determination and hatching success in the gopher tortoise (Gopherus polyphemus). RUSSELL L. BURKE, MICHAEL A. EWERT, JEFFREY B. McLEMORE, AND DALE R. JACKSON     86

Marine turtle nesting, nest predation, hatch frequency, and nesting seasonality on the Osa Peninsula, Costa Rica. DANA L. DRAKE     89

Nesting activity of the loggerhead turtle, Caretta caretta, on Fethiye Beach, Turkey, in 1994. IBRAHIM BARAN AND OGUZ TURKOZAN     93

Stomach contents of commercially harvested adult alligator snapping turtles, Macroclemys temminckii. KEVIN N. SLOAN, KURT A. BUHLMANN, AND JEFFREY E. LOVICH     96

Distribution of nesting sites of sea turtles in Okinawajima and adjacent islands of the central Ryukyus, Japan. AKIRA KIKUKAWA, NAOKI KAMEZAKI, KOICHI HIRATE, AND HIDETOSHI OTA     99

Mating behavior in captive alligator snapping turtles (Macroclemys temminckii). J. BRENT HARREL, NEIL H. DOUGLAS, MAURY M. HARDAWAY, AND R. DALE THOMAS     101

Meat on the move: diet of a predatory turtle, Deirochelys reticularia (Testudines: Emydidae). DALE R. JACKSON     105

Kemp’s ridley sea turtle (Lepidochelys kempii) tracked by satellite telemetry from Louisiana to nesting beach at Rancho Nuevo, Tamaulipas, Mexico. MAURICE L. RENAUD, JAMES A. CARPENTER, JO A. WILLIAMS, AND ANDRE M. LANDRY, JR.     108

Observations on Manouria impressa at Phu Luang Wildlife Sanctuary, northeastern Thailand. TANYA CHAN-ARD, KUMTHORN THIRAKHUPT, AND PETER PAUL VAN DIJK     109

Commentaries and Reviews

Possible demographic and ecologic consequences of sex ratio manipulation in turtles. JEFFREY E. LOVICH     114

On the absence of spur-thighed tortoises, Testudo graeca, from Egypt. JAMES R. BUSKIRK     118

Turtle Poetry

To a box turtle. JOHN UPDIKE     121

Newsnotes and Announcements 

Chelonian Research Foundation Linnaeus Fund: 1995 grant recipients. ANDERS G.J. RHODIN     122


Volume 2, Number 2 – October 1996

Special Focus Issue
The Leatherback Turtle, Dermochelys coriacea




Nesting Populations and Conservation

Leatherback turtles (Dermochelys coriacea) on St. Croix, U.S. Virgin Islands: fifteen years of conservation. RALF H. BOULON, JR., PETER H. DUTTON, AND DONNA L. McDONALD     141

Use of PIT tags and photoidentification to revise remigration estimates of leatherback turtles (Dermochelys coriacea) nesting in St. Croix, U.S. Virgin Islands, 1979–1995. DONNA L. McDONALD AND PETER H. DUTTON     148

Nesting of the leatherback turtle (Dermochelys coriacea) in Tongaland, KwaZulu-Natal, South Africa, 1963–1995. GEORGE R. HUGHES     153

Leatherback turtle, Dermochelys coriacea, nesting and nest success at Tortuguero, Costa Rica, in 1990–1991. ALISON J. LESLIE, DAVID N. PENICK, JAMES R. SPOTILA, AND FRANK V. PALADINO     159

Leatherback turtle, Dermochelys coriacea, nesting at Tortuguero, Costa Rica, in 1995. CATHI L. CAMPBELL, CYNTHIA J. LAGUEUX, AND JEANNE A MORTIMER     169


Biology and conservation of leatherback turtles, Dermochelys coriacea, at Playa Langosta, Costa Rica. ANNY CHAVES, GUSTAVO SERRANO, GUILLERMO MARIN, EUGENIA ARGUEDAS, ALEJANDRO JIMENEZ, AND JAMES R. SPOTILA     184

Subsistence hunting of leatherback turtles, Dermochelys coriacea, in the Kai Islands, Indonesia. ALEXIS SUAREZ AND CHRISTOPHER H. STARBIRD     190

Decline of the leatherback population in Terengganu, Malaysia, 1956–1995. ENG-HENG CHAN AND HOCK-CHARK LIEW     196

Leatherback turtles, Dermochelys coriacea, nesting in French Guiana, 1978–1995. MARC GIRONDOT AND JACQUES FRETEY     204

Worldwide population decline of Dermochelys coriacea: are leatherback turtles going extinct? JAMES R. SPOTILA, ARTHUR E. DUNHAM, ALISON J. LESLIE, ANTHONY C. STEYERMARK, PAMELA T. PLOTKIN, AND FRANK V. PALADINO     209

Biology and Morphology

Respiratory physiology of adult leatherback turtles (Dermochelys coriacea) while nesting on land. FRANK V. PALADINO, JAMES R. SPOTILA, MICHAEL P. O’CONNOR, AND ROBERT E. GATTEN, JR.     223

Reproductive physiology of nesting leatherback turtles (Dermochelys coriacea) at Las Baulas National Park, Costa Rica. DAVID C. ROSTAL, FRANK V. PALADINO, RHONDA M. PATTERSON, AND JAMES R. SPOTILA     230

Shallow water diving by leatherback turtles in the South China Sea. SCOTT A. ECKERT, HOCK-CHARK LIEW, KAREN L. ECKERT, AND ENG-HENG CHAN     237

Age and growth in leatherback turtles, Dermochelys coriacea (Testudines: Dermochelyidae): a skeletochronological analysis. GEORGE R. ZUG AND JAMES F. PARHAM     244

Electron microscopic analysis of vascular cartilage canals in the humeral epiphysis of hatchling leatherback turtles, Dermochelys coriacea. JOHANNES A.G. RHODIN, ANDERS G.J. RHODIN, AND JAMES R. SPOTILA     250

Systematics and Evolution

On the availability of the name Dermochelys coriacea schlegelii (Garman, 1884) as a species or subspecies of the leatherback turtle. LEO DANIEL BRONGERSMA     261

Evolution and phylogeny of the leatherback turtles (Dermochelyidae), with descriptions of new fossil taxa. ROGER CONANT WOOD, JONNIE JOHNSON-GOVE, EUGENE S. GAFFNEY, AND KEVIN F. MALEY     266


Books on Marine Fish, in which True Figures of the Fish are Presented. Book XVI. Chapters II–V. On Turtles. GUILLAUME RONDELET. [1554]. Translated and annotated from Latin by KENNETH F. KITCHELL, JR., arranged and edited by ANDERS G.J. RHODIN     287

Commentaries and Reviews

Are leatherbacks really threatened with extinction? PETER C.H. PRITCHARD     303

Turtle Poetry

Awakening. KEVIN GRIMES     306


Volume 2, Number 3 – April 1997




A new species of mud turtle (genus Kinosternon) from Jalisco and Colima, Mexico, with notes on its natural history. JAMES F. BERRY, MICHAEL E. SEIDEL, AND JOHN B. IVERSON     329

Life history and ecology of the Sonoran mud turtle (Kinosternon sonoriense) in southeastern Arizona: a preliminary report. RICHARD C. VAN LOBEN SELS, JUSTIN D. CONGDON, AND JOSIAH T. AUSTIN     338

Response of embryos of the red-eared turtle (Trachemys scripta elegans) to experimental exposure to water-saturated substrates. JOHN K. TUCKER, FREDRIC J. JANZEN, AND GARY L. PAUKSTIS     345

Growth, reproduction, and survivorship in the red-eared turtle, Trachemys scripta elegans, in Illinois, with conservation implications. JOHN K. TUCKER AND DON MOLL     352

Home range characteristics of radiotagged gopher tortoises on Kennedy Space Center, Florida. REBECCA B. SMITH, DAVID R. BREININGER, AND VICKIE L. LARSON     358

Ecology of the eastern spiny softshell, Apalone spinifera spinifera, in the Lamoille River, Vermont. TERRY E. GRAHAM AND ANDREW A. GRAHAM     363

Clutch size and frequency in Florida box turtles (Terrapene carolina bauri): implications for conservation. C. KENNETH DODD, JR.     370

A comparative spotting-scope study of the distribution and relative abundance of river cooters (Pseudemys concinna) in western Kentucky and southern Mississippi. PETER V. LINDEMAN     378

Redescription of the Arakan forest turtle Geoemyda depressa Anderson 1875 (Testudines: Bataguridae). JOHN B. IVERSON AND WILLIAM P. McCORD     384

Variation in shell morphology and color of Hermann’s tortoise, Testudo hermanni, in southern Europe. GHISLAINE GUYOT AND BERNARD DEVAUX     390

Young desert tortoise survival: energy, water, and food requirements in the field. KENNETH A. NAGY, DAVID J. MORAFKA, AND REBECCA A. YATES     396

Notes and Field Reports

The spotted turtle (Clemmys guttata) in central Florida. MARY E. BARNWELL, PETER A. MEYLAN, AND TIMOTHY WALSH     405

Radiographic determination of fecundity: is the technique safe for developing turtle embryos? THOMAS G. HINTON, PETER D. FLEDDERMAN, JEFFREY E. LOVICH, JUSTIN D. CONGDON, AND J. WHITFIELD GIBBONS     409

Maximum size of the Egyptian tortoise, Testudo kleinmanni. BALAZS L. FARKAS, LASZLO SASVARI, AND JAMES R. BUSKIRK     415

New locality records for freshwater turtles from the Yucatan Peninsula, Mexico. JAMES R. BUSKIRK     415

Problems for conservation of pond turtles (Emys orbicularis) in central Italy: is the introduced red-eared turtle (Trachemys scripta) a serious threat? LUCA LUISELLI, MASSIMO CAPULA, DARIO CAPIZZI, ERNESTO FILIPPI, VERONICA TRUJILLO JESUS, AND CLAUDIO ANIBALDI     417

Occurrence of double clutching in the southwestern pond turtle, Clemmys marmorata pallida, in the Los Angeles Basin. ROBERT H. GOODMAN, JR.     419

Allometry in the loggerhead turtle, Caretta caretta. NAOKl KAMEZAKl AND MASAFUMI MATSUI     421

Incidental capture of sea turtles in shrimp trawls with and without TEDs in U.S. Atlantic and Gulf waters. MAURICE L. RENAUD, JAMES M. NANCE, ELIZABETH SCOTT-DENTON, AND GREGG R. GITSCHLAG     425

Thermoregulation of the European pond turtle, Emys orbicularis, in central Italy. CORINNE DI TRANI AND MARCO A.L. ZUFFI     428

Dietary overlap in three sympatric congeneric freshwater turtles (Pseudemys) in Florida. KAREN A. BJORNDAL, ALAN B. BOLTEN, CYNTHIA J. LAGUEUX, AND DALE R. JACKSON     430

Predation by the imported fire ant (Solenopsis invicta) on loggerhead sea turtle (Caretta caretta) nests on Wassaw National Wildlife Refuge, Georgia. ROBERT A. MOULIS     433

Green turtle, Chelonia mydas, with fused carapacial scutes nesting at Atol das Rocas, Brazil. CLAUDIO BELLINI AND TAISI MARIA SANCHES     437

Commentaries and Reviews

Movement of hawksbill turtles — a different perspective on the DNA data. N. MROSOVSKY     438

Movement of hawksbill turtles: what scale is relevant to conservation, and what scale is resolvable with mtDNA data? BRIAN W. BOWEN AND ANNA L. BASS     440

Resolutions of the participants at the 17th Annual Symposium on Sea Turtle Biology and Conservation. Compiled by JEANETTE WYNEKEN AND DEBORAH T. CROUSE     442

Linnaeus Fund Research Reports

Seasonal movements and habitat preferences of spotted turtles (Clemmys guttata) in north central Connecticut. KATHLEEN M. PERILLO     445

Ecology and natural history of the wood turtle (Clemmys insculpta) in southern New Hampshire. SHEILA E. TUTTLE AND DAVID M. CARROLL     447

Geographic variation in the giant softshell turtle, Pelochelys bibroni. ROBERT G. WEBB     450

Orientation by the Australian eastern long-necked turtle, Chelodina longicollis. TERRY E. GRAHAM     450

Impact of egg content on post-hatching size, body composition, and performance in the common snapping turtle (Chelydra serpentina). MICHAEL S. FINKLER     452

Turtle Poetry

Ancient ones. GRACE S. McLAUGHLIN     456

Newsnotes and Announcements

Chelonian Research Foundation Linnaeus Fund: 1996 grant recipients. ANDERS G.J. RHODIN     457
The 17th Annual Sea Turtle Symposium — largest ever. JEANETTE WYNEKEN     457
El Mundo de la Tortuga: the leatherback turtle museum. CORINNE VIGNON, UBALDO ESTEBAN, LAURENT LOMBART, AND MAXIME MARQUEZ     458
The plight of the giant tortoises of Changuu Island, Zanzibar. STEVE TOLAN AND JOHN E. COOPER     459
Amphibian and Reptile Conservation — a new journal. CRAIG HASSAPAKIS     460
Three turtle conservation groups win Getty Prize. GILLIAN HAGGERTY     460
The 5th Annual Workshop on Natural History and Conservation of Florida Turtles. GEORGE L. HEINRICH     461


Volume 2, Number 4 – December 1997


Editorial comment. Anders G.J. Rhodin, Peter C.H. Pritchard, John L. Behler, and Russell A. Mittermeier          481


Population structure, activity, and sexual dimorphism in a central Florida population of box turtles, Terrapene carolina bauri. Melissa A. Pilgrim, Terence M. Farrell, and Peter G. May      483

Behavior, hibernacula, and thermal relations of softshell turtles (Trionyx spiniferus) overwintering in a small stream. Michael V. Plummer and J. Cary Burnley       489

Home range of the turtle, Phrynops rufipes, in an isolated reserve in central Amazônia, Brazil. William E. Magnusson, Aldeniza Cardoso de Lima, Valdenise Lopes da Costa, and Richard C. Vogt            494

Restoration of epidermal scute patterns during regeneration of the chelonian carapace. Gerald Kuchling          500

Patterns of size and longevity of gopher tortoise (Gopherus polyphemus) burrows: implications for the longleaf pine ecosystem. Craig Guyer and Sharon M. Hermann 507

Activity, habitat, and movement patterns of softshell turtles (Trionyx spiniferus) in a small stream. Michael V. Plummer, Nathan E. Mills, and Steve L. Allen      514

Population structure, activity, and conservation of the neotropical freshwater turtle, Hydromedusa maximiliani, in Brazil. Franco Leandro Souza and Augusto Shinya Abe        521

Osteological characters of the shell and humerus in hinged tortoises of the African genus Kinixys. Donald G. Broadley     526

Age and growth of wild Kemp’s ridley turtles (Lepidochelys kempi): cumulative results of tagging studies in Florida. Jeffrey R. Schmid and Wayne N. Witzell        532

Nesting physiology of Kemp’s ridley sea turtles, Lepidochelys kempi, at Rancho Nuevo, Tamaulipas, Mexico, with observations on population estimates. David C. Rostal, Janice S. Grumbles, Richard A. Byles, René Márquez-M., and David Wm. Owens           538

The phylogeny of cheloniid sea turtles revisited. James F. Parham and David E. Fastovsky       548

Mauremys pritchardi, a new batagurid turtle from Myanmar and Yunnan, China. William P. McCord      555

Notes and Field Reports

Differences in body size among populations of red-eared sliders (Trachemys scripta elegans) subjected to different levels of harvesting. Lisa M. Close and Richard A. Seigel       563

Ecological distribution and population dynamics of the Mediterranean tortoise, Testudo graeca nikolskii, in xerophytic forests of the western Caucasus, southern Russia. Alexander A. Inozemtsev        567

Delayed feeding in neonatal Kemp’s ridley, Lepidochelys kempii: a captive sea turtle management technique. Clark T. Fontaine and Theodore D. Williams        573

Growth of the turtle, Phrynops rufipes, in central Amazônia, Brazil. William E. Magnusson, Aldeniza Cardoso de Lima, Valdenise Lopes da Costa, and Odilon Pimentel de Lima           576

Estimating the time between hatching of sea turtles and their emergence from the nest. Matthew H. Godfrey and N. Mrosovsky         581

Predation upon olive ridley sea turtles (Lepidochelys olivacea) by the American crocodile (Crocodylus acutus) at Playa Nancite, Costa Rica. Rudy M. Ortiz, Pamela T. Plotkin, and David Wm. Owens           585

Growth and reproductive estimates from alligator snapping turtles, Macroclemys temminckii, taken by commercial harvest in Louisiana. Anton D. Tucker and Kevin N. Sloan      587

Record-sized tortoises, Testudo graeca ibera and Testudo hermanni boettgeri, from Bulgaria. Vladimir A. Beshkov    593

Reproductive biology and demography of gopher tortoises (Gopherus polyphemus) from the western portion of their range. Kelley R. Smith, Janie A. Hurley, and Richard A. Seigel       596

Decline of the loggerhead turtle, Caretta caretta, nesting on Senri Beach in Minabe, Wakayama, Japan. Katsufumi Sato, Takeharu Bando, Yoshimasa Matsuzawa, Hideji Tanaka, Wataru Sakamoto, Shingo Minamikawa, and Kiyoshi Goto  600

A comparison of plastral scute lengths among members of the box turtle genera Cuora and Terrapene. Carl H. Ernst, Jeffrey E. Lovich, Arndt F. Laemmerzahl, and Steve Sekscienski   603

An ecotourism initiative to increase awareness and protection of marine turtles in Brazil: the Turtle by Night Program. Claudia Figueiredo Vieitas and Maria Ângela Marcovaldi           607

An anomalous specimen of Pelusios sinuatus lacking mesoplastra. Donald G. Broadley     610

Characteristics of hibernacula use by spotted turtles, Clemmys guttata, in Ohio. Timothy L. Lewis and John Ritzenthaler         611

Observations of reproductive behavior of male green turtles (Chelonia mydas) at a nesting beach in Cyprus. Annette C. Broderick and Brendan J. Godley        615

Kachuga sylhetensis recorded from northern Bengal, with notes on turtles of Gorumara National Park, eastern India. Indraneil Das      616

Commentaries and Reviews

Galápagos tortoise nomenclature: still unresolved. George R. Zug   618

Galápagos tortoise nomenclature: a reply. Peter C.H. Pritchard       619

In Memoriam. A man to envy: James J. Parsons, 1915–1997. David R. Stoddart     621

Linnaeus Fund Research Reports

Molecular phylogeography of the western pond turtle (Clemmys marmorata): preliminary results. Fredric J. Janzen, Stacia L. Hoover, and H. Bradley Shaffer   623


Reptiles of the Autonomous Republic of Turkmenistan. Chary A. Ataev. [1985]. Translated from Russian by Balázs Farkas, arranged by James R. Buskirk     627

Turtle Poetry

Wake, Methuselah! (Tortoise in midwinter at the London Zoo). Author Unknown 635

Newsnotes and Announcements

Chelonian conservation news. Allen Salzberg            636
Seychelles Giant Tortoise Conservation Project        638
Sea Turtle Survival League      638
Reviewers for Chelonian Conservation and Biology Volume 2 (1996 – 1997)          639
International Conference on Turtles and Tortoises   640
18th International Symposium on Sea Turtle Biology and Conservation      640


Volume 3 (1998–2000)


Volume 3, Number 1 – August 1998


Taxonomy of Indian Ocean giant tortoises (Dipsochelys). Justin Gerlach and Laura Canning          3

Distribution and habitats of the Euphrates softshell turtle, Rafetus euphraticus, in southeastern Anatolia, Turkey, with observations on biology and factors endangering its survival. Ertan Taskavak and Mehmet K. Atatür           20

Temperature effect on incubation period in the yellow-spotted river turtle, Podocnemis unifilis, in the Colombian Amazon. Vivian P. Páez and Brian C. Bock   31

Impact of agricultural development on a population of wood turtles (Clemmys insculpta) in southern Québec, Canada. Raymond A. Saumure and J. Roger Bider  37

On growth and morphometrics of leopard tortoises, Geochelone pardalis, in Serengeti National Park, Tanzania, with observations on effects of bushfires and latitudinal variation in populations of eastern Africa. Michael R.K. Lambert, Kenneth L.I. Campbell, and Jonathan D. Kabigumila 46

Decline of the olive ridley sea turtle (Lepidochelys olivacea) nesting assemblage at Nancite Beach, Santa Rosa National Park, Costa Rica. Roldán A. Valverde, Stephen E. Cornelius, and Claudette L. Mo            58

Population structure and movement patterns of alligator snapping turtles (Macroclemys temminckii) in northeastern Arkansas. Stanley E. Trauth, J.D. Wilhide, and Anthony Holt          64

Water transport by nesting painted turtles (Chrysemys picta marginata) in Michigan. Owen M. Kinney, Roy D. Nagle, and Justin D. Congdon   71

Demography and population status of the flattened musk turtle, Sternotherus depressus, in the Black Warrior River Basin of Alabama. Karan A. Bailey and Craig Guyer           77

Notes and Field Reports

Assessment and initial treatment of cold-stunned sea turtles. Samuel S. Sadove, Robert Pisciotta, and Robert DiGiovanni      84

Ecological notes on feral populations of Trachemys scripta elegans in northern Taiwan. Tien-Hsi Chen and Kuang-Yang Lue      87

Testosterone titers of immature loggerhead sea turtles (Caretta caretta) incidentally caught in the central Mediterranean: a preliminary sex ratio study. Paolo Casale, Guido Gerosa, Roberto Argano, Salvatore Barbaro, and Gaetano Fontana     90

Female-biased sex ratio of juvenile loggerhead sea turtles in Georgia. C. Robert Shoop, Carol A. Ruckdeschel, and Robert D. Kenney    93

Reproductive ecology of the Indian softshell turtle, Aspideretes gangeticus, in northern India. Karthikeyan Vasudevan  96

Morphometrics of the fossil box turtle, Terrapene innoxia Hay 1916, from Florida. Carl H. Ernst, James C. Wilgenbusch, Timothy P. Boucher, and Steven W. Sekscienski 99

Use of passive integrated transponder (PIT) tags for marking small freshwater turtles. Kurt A. Buhlmann and Tracey D. Tuberville        102

First record of fibropapilloma on an olive ridley turtle in Nicaragua. Todd M. Steiner, Randall Arauz Vargas, and Pandora Martinez      105

Food habits of the pelomedusid turtle Pelusios castaneus castaneus in southeastern Nigeria. Luca Luiselli          106

Annual deposition of scute rings in the western pond turtle, Clemmys marmorata. R. Bruce Bury and David J. Germano            108

Bait preferences of southeastern United States Coastal Plain riverine turtles: fish or fowl? John B. Jensen           109

Sexual dimorphism of neonate eastern spiny softshells, Apalone spinifera spinifera. Terry E. Graham and Cynthia B. Cobb        111

Molecules, morphology, and mud turtle phylogenetics (family Kinosternidae). John B. Iverson     113

Commentaries and Reviews

How to minimize risk and optimize information gain in assessing reproductive condition and fecundity of live female chelonians. Gerald Kuchling            118

Age determination in turtles: evidence of annual deposition of scute rings. David J. Germano and R. Bruce Bury 123

Linnaeus Fund Research Reports

Identification of Seychelles giant tortoises. Justin Gerlach and Laura Canning        133

Current status and conservation of the river cooter (Pseudemys concinna) in southern Illinois. Michael J. Dreslik           135

Of deadwood and map turtles (Graptemys): an analysis of species status for five species in three river drainages using replicated spotting-scope counts of basking turtles. Peter V. Lindeman 137

Nesting ecology of the striped mud turtle (Kinosternon baurii) in central Florida. Dawn S. Wilson 142

Morphology and histology of the digestive system of the desert tortoise, Gopherus agassizi. Robert M. Winokur, Martin Hicks, and Naomi Chaney            144

Turtle Poetry

Turtle Soup. Marilyn Chin       146

Newsnotes and Announcements

Chelonian conservation news. Allen Salzberg            147
Chelonian Research Foundation Linnaeus Fund: 1997 grant recipients. Anders G.J. Rhodin           150
Turtle decoys – a request for information. René E. Honegger          151
The 18th Annual Sea Turtle Symposium – Student Paper Awards. Anders G.J. Rhodin and F. Alberto Abreu Grobois       152
Symposium on the Status and Conservation of Florida Turtles. George L. Heinrich 152


Volume 3, Number 2 – April 1999

Special Focus Issue
The Hawksbill Turtle, Eretmochelys imbricata


Editorial comment. Anders G.J. Rhodin and Peter C.H. Pritchard     171


Biology and Status of the Hawksbill in the Caribbean.
A Review Prepared by the IUCN/SSC Marine Turtle Specialist Group.

Foreword. Maria Angela (Neca) Marcovaldi  173

Introduction. Conservation of hawksbill sea turtles: perceptions and realities. Karen A. Bjorndal  174

Status of the hawksbill turtle (Eretmochelys imbricata) in the Caribbean region. Anne B. Meylan 177

Population modeling and implications for Caribbean hawksbill sea turtle management. Deborah T. Crouse         185

International movements of immature and adult hawksbill turtles (Eretmochelys imbricata) in the Caribbean region. Anne B. Meylan 189

Genetic analysis to elucidate the natural history and behavior of hawksbill turtles (Eretmochelys imbricata) in the wider Caribbean: a review and re-analysis. Anna L. Bass         195


Status justification for listing the hawksbill turtle (Eretmochelys imbricata) as Critically Endangered on the 1996 IUCN Red List of Threatened Animals.  Anne B. Meylan and Marydele Donnelly    200

Differential tag retention in Caribbean hawksbill turtles. Robert P. van Dam and Carlos E. Diez     225

Population structure of hawksbill turtles on a foraging ground in the Dominican Republic. Yolanda M. León and Carlos E. Diez  230

Internesting movements and behavior of hawksbill turtles (Eretmochelys imbricata) around Buck Island Reef National Monument, St. Croix, U.S. Virgin Islands. Christopher H. Starbird, Zandy Hillis-Starr, James T. Harvey, and Scott A. Eckert     237

Population ecology and demographic implications drawn from an 11-year study of nesting hawksbill turtles, Eretmochelys imbricata, at Jumby Bay, Long Island, Antigua, West Indies. James I. Richardson, Rebecca Bell, and Thelma H. Richardson      244

Estimating the annual size of hawksbill (Eretmochelys imbricata) nesting populations from mark-recapture studies: the use of long-term data to provide statistics for optimizing survey effort. Rhema Kerr, James I. Richardson, and Thelma H. Richardson        251

Reproduction and nesting of the hawksbill turtle, Eretmochelys imbricata, in the Cuban archipelago. Felix Moncada, Elvira Carrillo, Ana Saenz, and Gonzalo Nodarse 257

Hawksbill turtles (Eretmochelys imbricata) in Cuba: an assessment of the historical harvest and its impacts. Elvira Carrillo, Grahame J.W. Webb, and S. Charlie Manolis 264

Nesting of the hawksbill turtle, Eretmochelys imbricata, in Río Lagartos, Yucatán, Mexico, 1990–1997. Mauricio Garduño-Andrade      281

Increases in hawksbill turtle (Eretmochelys imbricata) nestings in the Yucatán Peninsula, Mexico, 1977–1996: data in support of successful conservation? Mauricio Garduño-Andrade, Vicente Guzmán, Emma Miranda, Raquel Briseño-Dueñas, and F. Alberto Abreu-Grobois   286

Genetic sourcing for the hawksbill turtle, Eretmochelys imbricata, in the northern Caribbean region. Rogelio Díaz-Fernández, Toshinao Okayama, Tomouki Uchiyama, Elvira Carrillo, Georgina Espinosa, René Márquez, Carlos Diez, and Hiroko Koike       296

Nesting and conservation management of hawksbill turtles (Eretmochelys imbricata) in northern Bahia, Brazil. Maria Angela Marcovaldi, Claudia F. Vieitas, and Matthew H. Godfrey   301

Juvenile Eretmochelys imbricata and Chelonia mydas in the Archipelago of Fernando de Noronha, Brazil. Taisi Maria Sanches and Claudio Bellini            308

The hawksbill turtle, Eretmochelys imbricata, in the Arabian Gulf. Nicolas J. Pilcher          312

Temporal distribution and periodicity in hawksbill turtles (Eretmochelys imbricata) nesting at Cousin Island, Republic of Seychelles, 1971–1997. Jeanne A. Mortimer and Roby Bresson          318

Hawksbill turtles, Eretmochelys imbricata, nesting on Redang Island, Terengganu, Malaysia, from 1993 to 1997. Eng-Heng Chan and Hock-Chark Liew     326

Reproductive biology of the hawksbill turtle, Eretmochelys imbricata, in Sabah, Malaysia. Nicolas J. Pilcher and Lamri Ali          330

Current status of nesting populations of the hawksbill turtle (Eretmochelys imbricata) in the Java Sea, Indonesia. Hiroyuki Suganuma, Naoki Kamezaki, and Akil Yusuf        337

Hawksbill turtle, Eretmochelys imbricata, nesting at Milman Island, northern Great Barrier Reef, Australia. Kirstin A. Dobbs, Jeffrey D. Miller, Colin J. Limpus, and André M. Landry, Jr.      344

Genetic diversity of the hawksbill turtle in the Indo-Pacific and Caribbean regions. Toshinao Okayama, Rogelio Díaz-Fernández, Yoshiyuki Baba, Matheus Halim, Osamu Abe, Naofumi Azeno, and Hiroko Koike      362

Turtle Poetry

Adelita sleeps…. Barbara Garrison     368
Caretta’s Return. Barbara Garrison    369

Newsnotes and Announcements

Chelonian conservation news. Allen Salzberg            370
Chelonian Research Foundation Linnaeus Fund: 1998 grant recipients. Anders G.J. Rhodin           373
The 19th Annual Sea Turtle Symposium – student presentation awards. Anders G.J. Rhodin and David Wm. Owens       374
2nd ASEAN Symposium and Workshop on Sea Turtle Biology and Conservation. Nicolas J. Pilcher 375
2nd International Symposium on Emys orbicularis. Alain Veysset and Barbara Livoreil       375
A Forum on the Biology and Conservation of Turtles in the Genus Clemmys. Sheila Tuttle 376
World’s first turtle shell stockpile to go up in flames as Miss World 1998 contestants look on. Jeanne A. Mortimer        376
New data reverse turtle’s spot on reptile tree. Carol Kaesuk Yoon   377
Sri Lanka herpetological publications available. Anslem de Silva      378
Chelonian Research Foundation announces a new turtle and tortoise newsletter and web site: www.chelonian.org. Anders G.J. Rhodin          378


Volume 3, Number 3 – December 1999


Editorial comment. Anders G.J. Rhodin         391


Home range and microhabitat use in the angonoka (Geochelone yniphora) in Madagascar. Lora L. Smith, Bourou Robert, Mahatoly Joby, and Sibo Clément          393

Environmental correlates of hatchling emergence in the red-eared turtle, Trachemys scripta elegans, in Illinois. John K. Tucker 401

Thermal profiles of sea turtle hatcheries and nesting areas at Praia do Forte, Brazil. Eugênia Naro-Maciel, N. Mrosovsky, and Maria Ângela Marcovaldi      407

Ecology and status of a wood turtle (Clemmys insculpta) population in West Virginia. Allan J. Niederberger and Michael E. Seidel         414

Nesting ecology of Podocnemis sextuberculata (Testudines, Pelomedusidae) in the Japurá River, Amazonas, Brazil. Juarez C.B. Pezzuti and Richard C. Vogt           419

Habitat selection and movements of diamondback terrapins, Malaclemys terrapin, in a Maryland estuary. Willem M. Roosenburg, Katherin L. Haley, and Scott McGuire       425

Home ranges of spotted turtles (Clemmys guttata) in southwestern Ohio. Timothy L. Lewis and Craig A. Faulhaber        430

Leopard tortoises (Geochelone pardalis) in Valley Bushveld, Eastern Cape, South Africa: specialist or generalist herbivores? Mervyn C. Mason, Graham I.H. Kerley, Craig A. Weatherby, and William R. Branch      435

Occurrence and trade of the golden turtle, Cuora trifasciata, in Laos. Robert J. Timmins and Khamkhoun Khounboline  441

Physical and microhabitat characteristics of burrows used by juvenile desert tortoises (Gopherus agassizii). Dawn S. Wilson, C. Richard Tracy, Kenneth A. Nagy, and David J. Morafka         448

Conservation genetics of the giant Amazon River turtle (Podocnemis expansa; Pelomedusidae) – inferences from two classes of molecular markers. Jack W. Sites, Jr., Nancy N. FitzSimmons, Nelson Jorge da Silva, Jr., and Vitor H. Cantarelli         454

Notes and Field Reports

Growth of slider turtles (Trachemys scripta) from temperate and tropical populations. John K. Tucker, Don Moll, and Edward O. Moll  464

Husbandry, behavior, and captive breeding of the Namaqualand speckled padloper (Homopus signatus signatus). Victor J.T. Loehr      468

Diet of Elseya purvisi and Elseya georgesi (Testudines: Chelidae), a sibling species pair of freshwater turtles from eastern Australia. Mathew Allanson and Arthur Georges 473

Short-term effectiveness of Passive Integrated Transponder (PIT) tags used in the study of Mediterranean marine turtles. Brendan J. Godley, Annette C. Broderick, and Sinead Moraghan 477

On the turtles of Hainan Island, southern China. Ron W.F. de Bruin and Harald G. Artner  479

Observations on predation of marine turtle nests at Akyatan, Turkey, eastern Mediterranean. Monica Aureggi, Guido Gerosa, and Sedat V. Yerli             487

Occurrence of the furrowed wood turtle (Rhinoclemmys areolata) on Turneffe Atoll, Belize. Steven G. Platt, William B. Karesh, John B. Thorbjarnarson, and Thomas R. Rainwater    490

Size differences in hind limbs and carapaces of hatchling green turtles (Chelonia mydas) from Hawaii and Florida, USA. Jeanette Wyneken, George H. Balazs, S.K.K. Murakawa, and Yvette Anderson    491

Feeding behavior and the saddleback shell of Dipsochelys arnoldi. Justin Gerlach  496

Long-term trends in size of stranded juvenile loggerhead sea turtles (Caretta caretta). C. Robert Shoop, Carol A. Ruckdeschel, and Robert D. Kenney            501

Effects of short-term water temperature variation on food consumption in painted turtles (Chrysemys picta marginata). Kristie D. Gianopulos and John W. Rowe 504

Habitat structures associated with juvenile gopher tortoise burrows on pine plantations in Alabama. Matthew J. Aresco           507

First record of the Nile soft-shelled turtle, Trionyx triunguis, from Kos Island, Greece, with comments on its occurrence in the eastern Mediterranean. Ertan Taskavak, Michael J. Reimann, and Wouter N. Polder          510

Cutaneous surface area in freshwater turtles. Paul A. Stone and John B. Iverson    512

Stomach contents of an olive ridley turtle (Lepidochelys olivacea) from the Gulf of Papua, Papua New Guinea. C. Sylvia Spring and Janet Gwyther            516

First record of introduced eastern Mediterranean tortoises, Testudo hermanni boettgeri, in southern France. Ghislaine Guyot and Peter C.H. Pritchard         518

Commentaries and Reviews

Estimating the time between hatching and emergence from the nest of sea turtles: effects of ignoring water potential. Anthony C. Steyermark            521

Linnaeus Fund Research Reports

Reproductive cycles of male and female giant tortoises (Geochelone nigra) on the Galápagos Islands by plasma steroid analysis and ultrasound scanning. Beatrix G. Schramm, Valentine A. Lance, and Miguel Casares      523

A test of the comparative influences of constant and fluctuating incubation temperatures on phenotypes of hatchling turtles. J. Sean Doody  529

Mercury concentration in the scutes of black sea turtles, Chelonia mydas agassizii, in the Gulf of California. Stephanie M. Presti, Antonio Resendiz S. Hidalgo, Albert E. Sollod, and Jeffrey A. Seminoff  531

Turtle Poetry

Ancient Chelonians. Anders G.J. Rhodin        534

Newsnotes and Announcements

Chelonian Research Foundation Linnaeus Fund: 1999 grant recipients. Anders G.J. Rhodin           535
A fish spearing decoy in the likeness of a Blanding’s turtle (Emydoidea blandingii). Philip A. Cochran       536
A new outlet for chelonian newsnotes and announcements: Turtle and Tortoise Newsletter. Heather J. Kalb and Allen Salzberg           536


Volume 3, Number 4 – November 2000

Special Focus Issue
The Blanding’s Turtle, Emydoidea blandingii


Editorial comment. Anders G.J. Rhodin         553

Special Focus Section


Blanding’s Turtle Workshop summary. John J. Moriarty       555


The Blanding’s turtles (Emydoidea blandingii) of Weaver Dunes, Minnesota. Michael J. Pappas, Bruce J. Brecke, and Justin D. Congdon           557

Nesting ecology and embryo mortality: implications for hatchling success and demography of Blanding’s turtles (Emydoidea blandingii). Justin D. Congdon, Roy D. Nagle, Owen M. Kinney, Matthew Osentoski, Harold W. Avery, Richard C. van Loben Sels, and Donald W. Tinkle        569

Population structure and reproductive ecology of Blanding’s turtle (Emydoidea blandingii) in Maine, near the northeastern edge of its range. Lisa A. Joyal, Mark McCollough, and Malcolm L. Hunter, Jr.        580

Spatial ecology of Blanding’s turtle in central Minnesota. Stephen A. Piepgras and Jeffrey W. Lang          589

Occurrence, habitat selection, and movement patterns of juvenile Blanding’s turtles (Emydoidea blandingii) in Kejimkujik National Park, Nova Scotia. Natalie L. McMaster and Thomas B. Herman 602

Movement of hatchling Blanding’s turtles (Emydoidea blandingii) in Nova Scotia in response to proximity to open water: a manipulative experiment. Jennifer A. McNeil, Thomas B. Herman, and K. Lorraine Standing     611

Growth and population structure of Emydoidea blandingii from western Nebraska. David J. Germano, R. Bruce Bury, and Mary Jennings         618

Thermal ecology of Blanding’s turtle in central Minnesota. Todd D. Sajwaj and Jeffrey W. Lang    626

Results of the nest protection program for Blanding’s turtle in Kejimkujik National Park, Canada: 1987–1997. K. Lorraine Standing, Thomas B. Herman, Maria Shallow, Terry Power, and Ian P. Morrison  637

Impact of a controlled wetland drawdown on Blanding’s turtles in Minnesota. Carol Dorff Hall and Francesca J. Cuthbert          643

Restoration of wetland and upland habitat for the Blanding’s turtle, Emydoidea blandingii. Erik Kiviat, Gretchen Stevens, Robert Brauman, Sven Hoeger, Peter J. Petokas, and Garrett G. Hollands    650

Notes and Field Reports

Predation of neonate Blanding’s turtles (Emydoidea blandingii) by short-tailed shrews (Blarina brevicauda). K. Lorraine Standing, Thomas B. Herman, and Ian P. Morrison       658

Developmental abnormalities in a northeastern population of Blanding’s turtle, Emydoidea blandingii. K. Lorraine Standing, Thomas B. Herman, and Ian P. Morrison    661

Commentaries and Reviews

Commercial exploitation of Blanding’s turtle, Emydoidea blandingii, and the wood turtle, Clemmys insculpta, for the live animal trade. John P. Levell   665


Regular Contributions


A new species of long-necked turtle (Testudines: Chelidae) from the Arnhem Land Plateau, Northern Territory, Australia. Scott Thomson, Rod Kennett , and Arthur Georges 675

Reproductive ecology of the loggerhead turtle, Caretta caretta, on Fethiye and Kizilot beaches, Turkey. Oguz Türkozan 686

Do wire cages protect sea turtles from foot traffic and mammalian predators? Melissa L. Mroziak, Michael Salmon, and Kirt Rusenko  693

Habitat selection and modification by the gopher tortoise, Gopherus polyphemus, in Georgia longleaf pine forest. Melissa D. Boglioli, William K. Michener, and Craig Guyer     699

Ultrasound scanning of ovaries and eggs in Galápagos tortoises, Geochelone nigra, on Santa Cruz Island, Galápagos. Beatrix G. Schramm, Miguel Casares, and Valentine A. Lance         706

Environmental characteristics of desert tortoise (Gopherus agassizii) burrow locations in an altered industrial landscape. Jeffrey E. Lovich and Ramona Daniels          714

Local harvest of hingeback tortoises, Kinixys erosa and K. homeana, in southwestern Cameroon. Dwight P. Lawson       722

Notes and Field Reports

The green turtle, Chelonia mydas, in the Saudi Arabian Gulf. Nicolas J. Pilcher       730

Use of a marked population of diamondback terrapins (Malaclemys terrapin) to determine impacts of recreational crab pots. Margaret E. Hoyle and J. Whitfield Gibbons        735

Update on permanent residency, persistence, and longevity in a 35-year study of a population of three-toed box turtles in Missouri. Elizabeth R. Schwartz         737

A translocation strategy for confiscated pancake tortoises. Luke James Harmon    738

Aquatic home ranges of female western pond turtles, Clemmys marmorata, at two sites in southern California. Robert H. Goodman, Jr. and Glenn R. Stewart       743

The identification of the holotype of Chelodina oblonga (Testudines: Chelidae) with a discussion of taxonomic implications. Scott Thomson    745

Observations of male green turtles (Chelonia mydas) on the nesting beach at Tortuguero National Park, Costa Rica. Sebastian Troëng 749

Predation of green (Chelonia mydas) and leatherback (Dermochelys coriacea) turtles by jaguars (Panthera onca) at Tortuguero National Park, Costa Rica. Sebastian Troëng 751

Mud accumulation in nesting aquatic turtles (Emydidae) in Illinois. John K. Tucker and Daniel A. Warner 753

Commentaries and Reviews

Sexing juvenile sea turtles: is there an accurate and practical method? Thane Wibbels, David Wm. Owens, and Colin J. Limpus 756

A response to Nicholas Mrosovsky’s Sustainable Use of Hawksbill Turtles: Contemporary Issues in Conservation. Peter C.H. Pritchard  761

Conservation and cultural diversity: reply to Pritchard. N. Mrosovsky         767

Turtle Poetry

Tragedy of the road. Don Blanding    770


Volume 4 (2001–2005)


Volume 4, Number 1 – September 2001


Editorial comment. Anders G.J. Rhodin         3


Observations on body size, sympatry, and niche divergence in softshell turtles (Trionychidae). Peter C.H. Pritchard        5

Sexing young free-ranging desert tortoises (Gopherus agassizii) using external morphology. James L. Boone and Eric A. Holt     28

The green turtle, Chelonia mydas, in Queensland, Australia: the Bramble Cay rookery in the 1979–1980 breeding season. Colin J. Limpus, David Carter, and Mark Hamann      34

Genetic population structure of two threatened South American river turtle species, Podocnemis expansa and Podocnemis unifilis. Brian C. Bock, Vivian P. Páez, and Matthew M. White          47

Morphometric model for sex assessment in hatchling olive ridley sea turtles. Jesús Emilio Michel-Morfin, Victor M. Gómez Muñoz, and Carmen Navarro Rodríguez      53

Deforestation and habitat loss for the Malagasy flat-tailed tortoise from 1963 through 1993. Stephen T. Tidd, John E. Pinder III, and Gary W. Ferguson  59

Demographic and ecological factors affecting conservation and management of the diamondback terrapin (Malaclemys terrapin) in South Carolina. J. Whitfield Gibbons, Jeffrey E. Lovich, Anton D. Tucker, Nancy N. FitzSimmons, and Judith L. Greene          66

Seasonal movements and reproduction in the striped mud turtle (Kinosternon baurii) from the southern Everglades, Florida. Walter E. Meshaka, Jr. and Eric Blind         75

Use of thermal springs for aquatic basking by the pig-nosed turtle, Carettochelys insculpta. J. Sean Doody, Rachel A. Sims, and Arthur Georges            81

A system-of-equations growth function for southern Great Barrier Reef green sea turtles. Milani Chaloupka      88

Some ecological parameters of the wood turtle, Clemmys insculpta, in southeastern Pennsylvania. Carl H. Ernst            94

Sexual dimorphism and morphological differentiation in European pond turtle (Emys orbicularis) populations from northwestern Spain. César Ayres Fernández and Adolfo Cordero Rivera 100

Migration routes and destination characteristics of post-nesting hawksbill turtles satellite-tracked from Barbados, West Indies. Julia A. Horrocks, Lotus A. Vermeer, Barry Krueger, Michael Coyne, Barbara A. Schroeder, and George H. Balazs     107

Terrestrial habitat use by aquatic turtles from a seasonally fluctuating wetland: implications for wetland conservation boundaries. Kurt A. Buhlmann and J. Whitfield Gibbons   115

Trace metal concentrations in blood of the Kemp’s ridley sea turtle (Lepidochelys kempii). Lisa O. Kenyon, André M. Landry, Jr., and Gary A. Gill            128

Habitat use and movements of radiotagged Blanding’s turtles (Emydoidea blandingii) in a suburban landscape. Cory S. Rubin, Richard E. Warner, and Daniel R. Ludwig  136

The loggerhead turtle, Caretta caretta, in Queensland: breeding migrations and fidelity to a warm temperate feeding area. Colin J. Limpus and Duncan J. Limpus        142

Notes and Field Reports

Notes on the distribution, life history, and exploitation of turtles in Sulawesi, Indonesia, with emphasis on Indotestudo forstenii and Leucocephalon yuwonoi. Steven G. Platt, Robert J. Lee, and Michael W. Klemens         154

Kyphosis in a musk turtle (Sternotherus odoratus) from Ontario, Canada. Raymond A. Saumure  159

Surface ultrastructure and retention of the egg caruncle in the European pond turtle, Emys orbicularis. Slawomir Mitrus          160

Response of juvenile softshell turtles (Apalone mutica) in a thermal gradient. Alan V. Nebeker and R. Bruce Bury           162

Husbandry, behavior, and captive breeding of the Nama padloper, Homopus bergeri, from southwestern Namibia. Alfred Schleicher and Victor J.T. Loehr         165

Mycoflora identified from failed green (Chelonia mydas) and loggerhead (Caretta caretta) sea turtle eggs at Heron Island, Australia. Andrea D. Phillott, C. John Parmenter, and Colin J. Limpus        170

A tortoise survey of Shwe Settaw Wildlife Sanctuary, Myanmar, with notes on the ecology of Geochelone platynota and Indotestudo elongata. Steven G. Platt, Saw Tun Khaing, Win Ko Ko, and Kalyar       172

Summertime foraging and feeding by immature loggerhead sea turtles (Caretta caretta) from Georgia. Michael G. Frick, Kristina L. Williams, and Lianne Pierrard           178

Winter survey of a gopher tortoise population in South Carolina. Tracey D. Tuberville and Michael E. Dorcas      182

Penis displays of snapping turtles (Chelydra serpentina) in response to handling: defensive or displacement behavior? Shane R. de Solla, Melinda Portelli, Holly Spiro, and Ronald J. Brooks     187

Maximum size and mass of the ploughshare tortoise, Geochelone yniphora. Miguel Pedrono and Tim Markwell 190

Nesting biology of the sea turtles of St. Kitts, West Indies. Joseph A. Butler            191

Assessment of sea turtle mortality rates in the Bahía Magdalena region, Baja California Sur, México. Susan C. Gardner and Wallace J. Nichols 197

Diet of the softshell turtle, Trionyx triunguis, in an Afrotropical forested region. Godfrey C. Akani, Dario Capizzi, and Luca Luiselli         200

Feeding of Dermochelys coriacea on medusae in the northwest Atlantic. Michael C. James and Thomas B. Herman       202

Over three decades of persistence of a small and apparently isolated population of painted turtles (Chrysemys picta) in a Kentucky reservoir. Peter V. Lindeman and A. Floyd Scott            206

On the possible occurrence of the marginated tortoise, Testudo marginata, in Turkey. Oguz Türkozan, Yusuf Kumlutas, and Çetin Ilgaz            208

Commentaries and Reviews

An overview of the North American turtle genus Clemmys Ritgen, 1828. Carl H. Ernst       211

Relative importance of thermal and nonthermal factors on the incubation period of sea turtle eggs. Matthew H. Godfrey and N. Mrosovsky   217

Linnaeus Fund Research Reports

Age estimation of Eretmochelys imbricata by schlerochronology of carapacial scutes. Anton D. Tucker, Damien Broderick, and Lorelei Kampe 219

Investigations of the ecology of Graptemys versa and Pseudemys texana in southcentral Texas. Peter V. Lindeman        223

Molecular systematics of emydine turtles. Chris R. Feldman and James Ford Parham        224

Genetic relatedness of populations of the mud turtle, Kinosternon baurii, from mainland Florida and the upper and Lower Florida Keys. Dawn S. Wilson and Stephen A. Karl    228

Turtle Poetry

Playa Grande beach. Michael H.J. Rhodin      231


Volume 4, Number 2 – December 2002

Special Focus Issue
The Gopherine Tortoises, Gopherus spp.


Editorial comment. Anders G.J. Rhodin         247


The Desert Tortoise, Gopherus agassizii

Defining the desert tortoise(s): our first priority for a coherent conservation strategy. Kristin H. Berry, David J. Morafka, and Robert W. Murphy            249

Is Gopherus agassizii a desert-adapted tortoise, or an exaptive opportunist? Implications for tortoise conservation. David J. Morafka and Kristin H. Berry           263

Reproduction in a desert tortoise (Gopherus agassizii) population on the Beaver Dam Slope, Washington County, Utah. Ann M. McLuckie and Rick A. Fridell          288

Reproduction of Gopherus agassizii in the Sonoran Desert, Arizona. Roy C. Averill-Murray            295

The annual reproductive cycle of the male and female desert tortoise: physiology and endocrinology. Valentine A. Lance and David C. Rostal 302

Chronology of sex determination in the desert tortoise (Gopherus agassizii). David C. Rostal, Thane Wibbels, Janice S. Grumbles, Valentine A. Lance, and James R. Spotila    313

Energy and water balance, diet, and reproduction of female desert tortoises (Gopherus agassizii). Brian Thomas Henen           319

Alien plants and fire in desert tortoise (Gopherus agassizii) habitat of the Mojave and Colorado deserts. Matthew L. Brooks and Todd C. Esque            330

Selective spring foraging by juvenile desert tortoises (Gopherus agassizii) in the Mojave Desert: evidence of an adaptive nutritional strategy. Olav T. Oftedal, Scott Hillard, and David J. Morafka          341

Diet selection by the desert tortoise in relation to the flowering phenology of ephemeral plants. W. Bryan Jennings      353

An experimental assessment of color, calcium, and insect dietary preferences of captive juvenile desert tortoises (Gopherus agassizii). Chris L. Okamoto         359

A landscape sampling protocol for estimating distribution and density patterns of desert tortoises at multiple spatial scales. Anthony J. Krzysik            366

Regional desert tortoise monitoring in the Upper Virgin River Recovery Unit, Washington County, Utah. Ann M. McLuckie, Deborah L. Harstad, Jesse W. Marr, and Rick A. Fridell      380

Spatial organization of desert tortoises and their burrows at a landscape scale. Jeffrey J. Duda, Anthony J. Krzysik, and Joel M. Meloche          387

Patterns of burrow use by desert tortoises (Gopherus agassizii) in southcentral Nevada. Kurt R. Rautenstrauch, Danny L. Rakestraw, Greg A. Brown, James L. Boone, and Patrick E. Lederle          398

Comparative dispersion of neonate and headstarted juvenile desert tortoises (Gopherus agassizii): a preliminary assessment of age effects. Lisa C. Hazard and David J. Morafka          406

Shell kinesis in juvenile desert tortoises, Gopherus agassizii. Marsha L. Marolda    410

Using growth ring counts to age juvenile desert tortoises (Gopherus agassizii) in the wild. Kristin H. Berry           416

A condition index for the desert tortoise (Gopherus agassizii). Kenneth A. Nagy, Brian T. Henen, Devesh B. Vyas, and Ian R. Wallis        425

Effects on survival of desert tortoises (Gopherus agassizii) urinating during handling. Roy C. Averill-Murray        430

Deaths of desert tortoises following periods of drought and research manipulation. Kristin H. Berry, E. Karen Spangenberg, Bruce L. Homer, and Elliott R. Jacobson       436

Impacts of vehicle road traffic on desert tortoise populations with consideration of conservation of tortoise habitat in southern Nevada. Karin von Seckendorff Hoff and Ronald William Marlow    449

Comparison of desert tortoise (Gopherus agassizii) populations in an unused and off-road vehicle area in the Mojave Desert. R. Bruce Bury and Roger A. Luckenbach  457

The Gopher Tortoise, Gopherus polyphemus

Reserve area requirements for gopher tortoises (Gopherus polyphemus). Jeannine Ott Eubanks, Jeff W. Hollister, Craig Guyer, and William K. Michener        464

A comparison of GIS and survey estimates of gopher tortoise habitat and numbers of individuals in Florida. Earl D. McCoy, Beth Stys, and Henry R. Mushinsky   472

Population biology of the gopher tortoise (Gopherus polyphemus) in southeast Georgia. David C. Rostal and Douglas N. Jones, Jr.       479

The Texas Tortoise, Gopherus berlandieri

Home range and dispersal of Texas tortoises, Gopherus berlandieri, in a managed thornscrub ecosystem. Richard T. Kazmaier, Eric C. Hellgren, and Donald C. Ruthven, III      488

Mycoplasmal Disease in Gopherus spp.

Application of diagnostic tests for mycoplasmal infections of desert and gopher tortoises, with management recommendations. Daniel R. Brown, Isabella M. Schumacher, Grace S. McLaughlin, Lori D. Wendland, Mary B. Brown, Paul A. Klein, and Elliott R. Jacobson  497

Turtle Poetry

Population density studies. Marian L. Griffey            508


Volume 4, Number 3 – December 2003


Editorial comment. Anders G.J. Rhodin         521


Reproductive biology and conservation status of the loggerhead sea turtle (Caretta caretta) in Espírito Santo State, Brazil. Cecília Baptistotte, João C.A. Thomé, and Karen A. Bjorndal        523

Post-nesting movements and behavior of loggerhead sea turtles (Caretta caretta) departing from east-central Florida nesting beaches. C. Kenneth Dodd, Jr. and Richard Byles        530

Previously unrecognized original type specimens of American turtles collected by John Le Conte in 1828. Roger Bour    537

Emergence success of flatback sea turtles (Natator depressus) at Fog Bay, Northern Territory, Australia. Sean J. Blamires and Michael L. Guinea            548

Influence of nest site selection on predation of flatback sea turtle (Natator depressus) eggs by varanid lizards in northern Australia. Sean J. Blamires, Michael L. Guinea, and Robert I.T. Prince  557

Diagnostic characteristics in lower Tennessee River populations of the map turtles Graptemys pseudogeographica and Graptemys ouachitensis. Peter V. Lindeman      564

Activity and thermal ecology of the box turtle, Terrapene ornata, at its southwestern range limit in Arizona. Michael V. Plummer        569

Daily and seasonal activity patterns and movements of juvenile box turtles (Terrapene carolina bauri) on Egmont Key, Florida. Alison Hamilton Jennings          578

Nesting and conservation of the hawksbill turtle, Eretmochelys imbricata, in the Pearl Cays, Nicaragua. Cynthia J. Lagueux, Cathi L. Campbell, and William A. McCoy       588

Comparative study of the nesting behavior of the green turtle, Chelonia mydas, during high- and low-density nesting periods at Ras Al-Hadd Reserve, Oman. A.Y.A. AlKindi, I.Y. Mahmoud, H.M. Al-Gheilani, C.S. Bakheit, A.A. Al-Habsi, and A. Al-Kiyumi          603

Status of the flatback sea turtle (Natator depressus) rookery on Crab Island, Australia, with notes on predation by crocodiles. Ronald W. Sutherland and Elizabeth G. Sutherland       612

Marine turtle conservation on Holbox Island, Quintana Roo, Mexico: the fishermen’s perceptions. Sonja Macys and George N. Wallace          620

Influence of geological factors on reproductive aspects of Podocnemis unifilis (Testudines, Pelomedusidae) on the Javaés River, Araguaia National Park, Brazil. Paulo Dias Ferreira Júnior, Adriana Malvasio, and Odimar Santos Guimarães   626

Morphological variation in the endangered Alabama red-bellied cooter (Pseudemys alabamensis) and taxonomic status of a population in Mississippi. Christopher J. Leary, James L. Dobie, Thomas M. Mann, and Peter S. Floyd   635

Biological prioritization of Asian countries for turtle conservation. Bryan L. Stuart and John Thorbjarnarson        642

Status of Testudo kleinmanni and T. werneri in Egypt. Sherif M. Baha El Din, Omar Attum, and Mindy Baha El Din          648

Home range and seasonal movements of the wood turtle (Glyptemys insculpta) in southern New Hampshire. Sheila E. Tuttle and David M. Carroll            656

Reproductive ecology of Podocnemis erythrocephala (Testudines: Podocnemididae) in the lower Inírida River, Colombia. Olga Victoria Castaño-Mora, Pedro Alberto Galvis-Peñuela, and Juan Gabriel Molano       664

Notes and Field Reports

Akinesis and plastral scute homologies in Sternotherus (Testudines: Kinosternidae). Peter C.H. Pritchard            671

Marine turtle nesting activity at Playa Naranjo, Santa Rosa National Park, Costa Rica, for the 1998–1999 season. Dana L. Drake, Jocelyn E. Behm, Meghan A. Hagerty, Philippe A. Mayor, Seth J. Goldenberg, and James R. Spotila  675

Ecology and conservation status of the Arakan forest turtle, Heosemys depressa, in western Myanmar. Steven G. Platt, Win Ko Ko, Kalyar, Myo Myo, Lay Lay Khaing, and Thomas R. Rainwater        678

Comparative ultrastructural carapace morphology in three freshwater turtles. David A. Wooten 682

Diet of sea turtles in southern Brazil. Leandro Bugoni, Lígia Krause, and Maria Virgínia Petry        685

Radiotransmitter attachment technique for box turtles (Terrapene spp.). Karen A. Wilson, Paul M. Cavanagh, and Jeffrey Villepique    688

Rediscovery of the critically endangered river terrapin, Batagur baska, in Cambodia, with notes on occurrence, reproduction, and conservation status. Steven G. Platt, Bryan L. Stuart, Heng Sovannara, Long Kheng, Kalyar, and Heng Kimchhay           691

Effects of sand hardness and human beach use on emergence success of loggerhead sea turtles on Yakushima Island, Japan. Hiromi Kudo, Akinobu Murakami, and Satomi Watanabe   695

Reproductive cycle of male snapping turtles (Chelydra serpentina) in southeastern Virginia. Lauren L. Glesenkamp, George R. Zug, and Joseph C. Mitchell           697

Natural hybridization between a Kemp’s ridley (Lepidochelys kempii) and loggerhead sea turtle (Caretta caretta) confirmed by molecular analysis. Robert C. Barber, Clark T. Fontaine, Joseph P. Flanagan, and Edward E. Louis, Jr.         701

Double-chambered nest cavities in the leatherback turtle. Stephanie Kamel and N. Mrosovsky    705

Summer movements and home range of the cooter turtle, Pseudemys concinna, in Illinois. Michael J. Dreslik, Andrew R. Kuhns, Christopher A. Phillips, and Benjamin C. Jellen          706

Comparative efficiency of different sampling techniques to obtain DNA from freshwater turtles. Nathalie Tessier and François-Joseph Lapointe            710

Effect of translocation on egg viability of the giant Amazon river turtle, Podocnemis expansa. Kelly Bonach, Maristela P. Miranda-Vilela, Marcelo C. Alves, and Luciano M. Verdade      712

First record of a gravid marine turtle from Chile. Angélica González, Leyla Miranda, and Juan Carlos Ortiz           716

Repetitive data-logger attachments to sea turtles using a new quick-release method. Sandra Storch and Solvin Zankl    717

Preliminary observations on reproductive parameters of the Sulawesi forest turtle (Leucocephalon yuwonoi) in captivity. Charles J. Innis        720

Population ecology of musk turtles (Sternotherus odoratus) in a lake in Virginia. Alison M. Holinka, Radhika Dave, and Randolph M. Chambers            721

Axial bifurcation in a bicephalic Chelonia mydas embryo. Cheong H. Diong, Lenny K.A. Tan, and Charles M.U. Leh          725

Commentaries and Reviews

Turtle management as scientific experimentation. William E. Magnusson  728

Linnaeus Fund Research Reports

Distribution, status, and biology of the radiated tortoise, Geochelone radiata, in southwest Madagascar. Thomas E.J. Leuteritz and Rollande Ravolanaivo    730

Domestic trade of tortoises and freshwater turtles in Cambodia. Rohan H.P. Holloway     733

Assessing the impact of human hunting activities on populations of forest tortoises (genus Kinixys) in the Niger Delta, Nigeria. Luca Luiselli    735

Interpond movement patterns of Chrysemys picta picta (eastern painted turtles) in Virginia. David R. Bowne     739

Turtle Poetry

The egg and the machine. Robert Frost         742


Volume 4, Number 4 – April 2005

Special Focus Issue
The Kemp’s Ridley Sea Turtle, Lepidochelys kempii


Editorial comment. Anders G.J. Rhodin         757

Special Focus Section


Introduction to the Kemp’s ridley focus issue. Peter C.H. Pritchard and David W. Owens   759


Status of the Kemp’s ridley sea turtle, Lepidochelys kempii. René Márquez-M., Patrick M. Burchfield, Juan Díaz-F., Manuel Sánchez-P., Miguel Carrasco-A., Carmen Jiménez-Q., Alma Leo-P., Rafael Bravo-G., and Jaime Peña-V.            761

A population model to estimate recovery time, population size, and management impacts on Kemp’s ridley sea turtles. Selina S. Heppell, Deborah T. Crouse, Larry B. Crowder, Sheryan P. Epperly, Wendy Gabriel, Terry Henwood, René Márquez, and Nancy B. Thompson 767

Time-series analyses of the relationship between nesting frequency of the Kemp’s ridley sea turtle and meteorological conditions. María del Carmen Jiménez-Quiroz, Anatoly Filonov, Irina Tereshchenko, and René Márquez-Millán 774

Reproductive parameters of the Kemp’s ridley sea turtle (Lepidochelys kempii) at Rancho Nuevo, Tamaulipas, Mexico. Wayne N. Witzell, Alfredo Salgado-Quintero, and Manuel Garduño-Dionte       781

Seasonal reproductive biology of the Kemp’s ridley sea turtle (Lepidochelys kempii): comparison of captive and wild populations. David C. Rostal            788

Population characteristics of Kemp’s ridley sea turtles in nearshore waters of the upper Texas and Louisiana coasts. André M. Landry, Jr., David T. Costa, F. Leonard Kenyon, II, and Michael S. Coyne  801

Kemp’s ridley sea turtle movements and migrations. Maurice L. Renaud and Jo A. Williams          808

Movements and home ranges of adult male Kemp’s ridley sea turtles (Lepidochelys kempii) in the Gulf of Mexico investigated by satellite telemetry. Donna J. Shaver, Barbara A. Schroeder, Richard A. Byles, Patrick M. Burchfield, Jaime Peña, René Márquez, and Hector J. Martinez           817

Developmental biology and ecology of the Kemp’s ridley sea turtle, Lepidochelys kempii, in the eastern Gulf of Mexico. Jeffrey R. Schmid and William J. Barichivich  828

Texans, turtles, and the early Kemp’s ridley population restoration project, 1963–67. Patrick M. Burchfield        835

Head-starting the Kemp’s ridley sea turtle, Lepidochelys kempii, at the NMFS Galveston Laboratory, 1978–1992: a review. Clark Fontaine and Donna Shaver 838

Analysis of the Kemp’s ridley imprinting and headstart project at Padre Island National Seashore, Texas, 1978–88, with subsequent nesting and stranding records on the Texas coast. Donna J. Shaver         846

On the sex ratio of juvenile Lepidochelys kempii in Georgia. Carol Ruckdeschel, C. Robert Shoop, and Robert D. Kenney            860

Diet analysis of Kemp’s ridley sea turtles (Lepidochelys kempii) in Virginia. Erin E. Seney and John A. Musick      864

Western North Atlantic waters: crucial developmental habitat for Kemp’s ridley and loggerhead sea turtles. Stephen J. Morreale and Edward A. Standora         872

Climatic and oceanographic factors affecting daily patterns of juvenile sea turtle cold-stunning in Cape Cod Bay, Massachusetts. Brett M. Still, Curtice R. Griffin, and Robert Prescott           883

Regular Contributions


Assessment of the threatened status of Testudo kleinmanni Lortet, 1883 (Testudines: Testudinidae) for the IUCN Red List. Jarmo Perälä         891

Conserving sea turtles in Canada: successful community-based collaboration between fishers and scientists. Kathleen Martin and Michael C. James 899

Ecology of the Jamaican slider turtle (Trachemys terrapen), with implications for conservation and management. Tracey D. Tuberville, Kurt A. Buhlmann, Rhema Kerr Bjorkland, and Doug Booher            908

Nesting activity and reproductive output of loggerhead sea turtles, Caretta caretta, over 19 seasons (1984–2002) at Laganas Bay, Zakynthos, Greece: the largest rookery in the Mediterranean. Dimitris Margaritoulis   916

Documenting the value of volunteer effort for sea turtle conservation in South Carolina. Sally R. Hopkins-Murphy and Joan S. Seithel  930

Notes and Field Reports

Bifid tail regeneration in a turtle, Emydura sp. (Testudines: Chelidae). Gerald Kuchling      935

Thermoregulation in captive Indian Ocean giant tortoises. Justin Gerlach   937

Noteworthy records and exploitation of chelonians from the Ayeyarwady, Chindwin, and Dokhtawady rivers, Myanmar. Steven G. Platt, Win Ko Ko, Lay Lay Khaing, Khin Myo Myo, Tint Lwin, Thanda Swe, Kalyar, and Thomas R. Rainwater       942

Distribution, natural history, and exploitation of Leucocephalon yuwonoi in central Sulawesi, Indonesia. Cris Hagen and Iwalani Y.S. Ching      948

Genetic analysis of mitochondrial DNA variation in eastern and western African spurred tortoises, Geochelone sulcata. Barbara Livoreil and Antoinette C. van der Kuyl          951

Distribution of the black-breasted leaf turtle (Geoemyda spengleri) on Hainan Island, China. Shi Haitao  954

Occurrence of diamondback terrapins, Malaclemys terrapin, on Bermuda: native or introduced? John Davenport, Anne F. Glasspool, and Lisa Kitson  956

Turtle Poetry

Turtles. Author Unknown       960


Volumes 5–19 (2006–2020)

Available at www.chelonianjournals.org